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Gehyra arnhemica OLIVER, PRASETYA, TEDESCHI, FENKER, ELLIS, DOUGHTY & MORITZ, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesEast Arnhem Land Gehyra 
SynonymGehyra arnhemica OLIVER, PRASETYA, TEDESCHI, FENKER, ELLIS, DOUGHTY & MORITZ 2020 
DistributionAustralia (Northern Territory)

Type locality: Long Billabong, Savannah Way, 2 km N Cox River, Roper Gulf, NT (−15.3067S, 135.3408E)  
Reproduction 
TypesHolotype. NTM R22626, adult male, collected by P. Horner, 19 May 1996.
Paratypes (N = 12). Northern Territory: NTM R38171 (field # CCM2271), NTM R38172 (field # CCM2272), Lake Katherine, NT (−14.3079, 135.0610E); NTM R38173 (field # CCM2408), Ngukurr, NT (−14.6550S, 134.7812E); NTM R38174 (field # CCM2573), NTM R38175 (field # CCM2575), NTM R38176 (field # CCM2576), Cox River area, Limmen NP, NT (−15.3167S, 135.3409E); NTM R38178 (field # CCM6511), NTM R38179 (field # CCM6516), Emu Springs, NT (−13.1562S, 134.8506E); NTM R38180 (field # CCM6554), NTM R38181 (field # CCM6555), NTM R38182 (field # CCM6595), 3.0 km SW Gikal, NT (−12.0813S, 136.2942E); NTM R38183 (field # CCM6647), 1.5 km NE Gikal, NT (−12.0522S, 136.3210E). 
DiagnosisDiagnosis. A large Gehyra species (up to 68.2 mm SVL) differing from all other Gehyra species outside of the G. australis complex as per the complex diagnosis above. Differs from other members of the G. australis complex in the combination of: moderate size within complex (adult SVL up to 68.2 mm, mean 63.0 mm); pre-cloacal pores in males numerous (21–26), often extending onto limbs and reduced in size distally; suture between first and second chin shields usually straight; second chin shields approximately two thirds length of first chin shields (mean ratio 0.67, range 0.57–0.77); and base colouration of adults smoky grey to brown, often with distinct and extensive pattern of darker-brown vermiculations, stripes, scalloping and/or blotches across the head, body and tail. Further diagnosed from other species within the G. australis complex genetically by three unique amino acids in the ND2 locus (Table 1). Some specimens, especially juveniles and females, may only be diagnosable from other members of the G. australis complex and G. chimera sp. nov. on the basis of locality and/or genetic data. Gehyra arnhemica sp. nov. overlaps or contacts the distribution of G. australis and G. gemina sp. nov. and is very similar morphologically to both species; however, males of G. arnhemica sp. nov. differ from both in having a higher number of pre-cloacal pores (21–26 vs. 11–14 in G. australis and 10–16 in G. gemina sp. nov.). It further differs from G. gemina sp. nov. in outer edge of first pair of chin shields in contact with second pair usually strait (vs. usually convex). Gehyra arnhemica sp. nov. further tends to differ from G. australis in having a more extensive and bolder pattern of dark brown blotches, lines and/or scalloping on the head, torso and tail. From G. lauta sp. nov., it differs in its smaller size (mean and maximum adult SVL, respectively: 68.2 mm and 63.0 mm vs. 71.4 mm and 83.1 mm). Gehyra arnhemica sp. nov. also overlap or abuts with the distribution of the rock-dwelling taxa G. borroloola and G. pamela; however, it can be readily differentiated from both by the absence of prominent pale spots and ocelli (vs. present), and by having smaller chin shields (extending to approximately level with posterior edge of second infralabial vs. approximately level with posterior edge of third infralabial) (see King, 1982). 
Comment 
EtymologyThe species epithet refers to the Arnhem Land region of the north-east Top End of the NT, in reference to the species occurrence and apparent endemism to the region. 
References
  • Oliver PM, Prasetya AM, Tedeschi LG, Fenker J, Ellis RJ, Doughty P, Moritz C. 2020. Crypsis and convergence: integrative taxonomic revision of the Gehyra australis group (Squamata: Gekkonidae) from northern Australia. PeerJ 8:e7971 - get paper here
 
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