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Gekko kuhli (STEJNEGER, 1902)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Kuhl's Flying Gecko, Gliding gecko, Kuhl’s Parachute Gecko
G: Kuhls Faltengecko 
SynonymLacerta homalocephala CREVELDT 1809 (pre-occupied)
Lacerta homalocephala — CUVIER 1831: 145
Platydactylus homalocephalus — DUMÉRIL & BIBRON 1836: 339
Ptychozoon homalocephalum — BOULENGER 1893: 316
Ptychozoon kuhli STEJNEGER 1902 (nom. nov.)
Ptychozoon homalocephalum — GÜNTHER 1864
Ptychozoon homalocephalum — BOULENGER 1885: 190
Ptychozoon homalocephalum — WEBER 1890: 165
Ptychozoon homalocephalum — DE ROOIJ 1915: 59
Ptychozoon kuhli — TAYLOR 1963: 810
Ptychozoon kuhli — KLUGE 1993
Ptychozoon kuhlii — COX et al. 1998: 81
Ptychozoon kuhli — CHAN-ARD et al. 1999: 131
Ptychozoon kulhli [sic] — PIANKA & VITT 2003: 181
Ptychozoon kuhli — NUR-AMALINA et al. 2017
Gekko (Ptychozoon) kuhli — WOOD et al. 2019
Gekko (Ptychozoon) kuhli — WOOD et al. 2020
Gekko cf. kuhli — FIGUEROA et al. 2023 
DistributionS Thailand (Nakhon Si Thammarat, Satun, Pattani), Myanmar (= Burma), Malaysian Peninsula (incl. Pulau Pangkor, Pulau Tioman, Johor: Pulau Besar), Nicobar Islands, Indonesia (Sumatra, Java, Borneo: Sabah, Sarawak; Simalur, Sulawesi, Bali, Natuna islands), Singapore

Type locality: not specified in original description (fide WERMUTH 1965, DAS & VIJAYAKUMAR 2009).  
Reproductionoviparous 
TypesHolotype: lost fide DE LISLE et al. 2013: 219 
DiagnosisDiagnosis (genus): Members of the genus are diagnosable from other Asian gekkonid genera (except Luperosaurus), in showing the following characters: medium-sized to large (SVL to 107.8 mm); cutaneous folds on limbs and on sides of body; digits dilated and strongly webbed, with undivided transverse lamellae; terminal phalanges united with dilated portion to its distal end and extending beyond; inner digit clawless; dorsal scales small, typically intermixed with large tubercles; pupil vertical and males with preanal and femoral pores (Brown et al. 1997).


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CommentSynonymy: Lacerta homalocephala CREVELDT 1809 is preoccupied by Lacerta homalocephala SUCHOW (or SUCKOW) 1798.

Type species: Lacerta homalocephala CREVELDT 1809 is the type species of the genus Ptychozoon KUHL & VAN HASSELT 1822. Lacerta homalocephala is also the type species of the subgenus Ptychozoon (Wood et al. 2019, 2020).

Comparisons: Grismer et al. 2019: 169 (Table 5) present a comparison of most species of Ptychozoon.

P. kuhli is highly variable and may consist of between five and nine unrecognized, distinct species (BROWN et al. 2013).

Distribution: not in India fide A.M. Bauer (pers. comm., 19 March 2015). 
EtymologyNamed after Dr. Heinrich Kuhl (1797-1821), a German ornithologist who was an assistant to Conrad Jacob Temminck. He also worked with Johan Conrad van Hasselt in Java before dying in Buitenzorg of a tropical disease.

See WERMUTH (1965: 154) and KLAVER (2007) for a discussion of the origin of the generic name Ptychozoon. 
References
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  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger, G.A. 1893. Viaggio di Leonardo Fea in Birmania e Regioni Vicine LII. Concluding report on the reptiles and batrachians obtained in Burma by Signor L. Fea, dealing with the collection made in Pegu and the Karin Hills in 1887-88. Annali Mus. civ. Genova (2) 13: 304-347 - get paper here
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  • Brown, Rafe M., John W. Ferner and Arvin C. Diesmos. 1997. Definition of the Philippine Parachute Gecko, Ptychozoon intermedium Taylor 1915 (Reptilia: Squamata: gekkonidae): Redescription, designation of a neotype, and comparisons with related species. Herpetologica 53 (3): 357-373. - get paper here
  • Brown, Rafe M.; Cameron D. Siler, L. Lee Grismer, Indraneil Das, Jimmy A. McGuire 2013. Phylogeny and cryptic diversification in Southeast Asian flying geckos. Molecular Phylogenetics and Evolution 66 (2):351-361 [2012] - get paper here
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