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Gekko rhacophorus (BOULENGER, 1899)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Sabah Flying Gecko 
SynonymGecko rhacophorus BOULENGER 1899: 451
Gecko rhacophorus — DE ROOIJ 1915: 55
Ptychozoon rhacophorus — KLUGE 1993
Ptychozoon rhacophorus — MANTHEY & GROSSMANN 1997: 248
Gekko rhacophorus — BROWN et al. 2012 (preliminary)
Ptychozoon rhacophorus — RÖSLER 2018: 22
Gekko (Rhacogekko) rhacophorus — WOOD et al. 2019 
DistributionMalaysia (Borneo, Sabah, Mount Kinabalu 700 m)

Type locality: Kadamaian River, Kina Balu, elevation: 2100 feet, North Borneo.  
Reproductionoviparous 
TypesHolotype: BMNH 1946.8.25.97 
DiagnosisDefinition (Subgenus Rhacogekko): Rhacogekko is a maximum crown-lineage name referring to Gekko (R.) rhacophorus originating as the sister lineage to Sundagekko and Pseudosundagekko. Gekko (R.) rhacophorus. Although unambiguous synapomorphies for the new subgenus have not been identified, Gekko (R.) rhacophorus differs from other subgenera by lacking lateral skin folds on head (infra-auricular cutaneous flaps), lacking imbricate dorsal parachute support scales and an enlarged terminal tail flap (Brown et al., 1997), and by having fully half-webbed fingers and toes (Boulenger, 1899; Malkamus et al., 2002, Wood et al. 2019). 
CommentPhylogeny: Based on DNA sequences, Ptychozoon rhacophorus is nested within Gekko (BROWN et al. 2012), and hence may have to be renamed.

Type species: Gekko (Rhacogekko) rhacophorus (Boulenger, 1899) is the type species of the subgenus Rhacogekko Wood et al. 2019. 
Etymology 
References
  • Boulenger, G A. 1899. Description of three new reptiles and a new batrachian from Mt. Kina Balu, North Borneo. Ann. Mag. Nat. Hist. (7) 4: 451 - 453 - get paper here
  • Brown, Rafe M. 1999. New species of parachute gecko (Squamata: Gekkonidae: genus Ptychozoon) from northeastern Thailand and central Vietnam. Copeia 1999 (4): 990-1001 - get paper here
  • Brown, Rafe M., John W. Ferner and Arvin C. Diesmos. 1997. Definition of the Philippine Parachute Gecko, Ptychozoon intermedium Taylor 1915 (Reptilia: Squamata: gekkonidae): Redescription, designation of a neotype, and comparisons with related species. Herpetologica 53 (3): 357-373. - get paper here
  • Brown, Rafe M.; Cameron D. Siler, L. Lee Grismer, Indraneil Das, Jimmy A. McGuire 2013. Phylogeny and cryptic diversification in Southeast Asian flying geckos. Molecular Phylogenetics and Evolution 66 (2):351-361 - get paper here
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das, I. & Vijayakumar, S.P. 2009. New species of Ptychozoon (Sauria: Gekkonidae) from the Nicobar Archipelago, Indian Ocean. Zootaxa 2095: 8–20 - get paper here
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp.
  • GRISMER, L. LEE; PERRY L. JR. WOOD, MYINT KYAW THURA, MARTA S. GRISMER, RAFE M. BROWN, BRYAN L. STUART 2018. Geographically structured genetic variation in Ptychozoon lionotum (Squamata: Gekkonidae) and a new species from an isolated volcano in Myanmar. Zootaxa 4514 (2): 202–214 - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Manthey, U. 1982. Die Gattung Ptychozoon, Teil 2 - Mit einem Bestimmungsschlüssel für die fünf Arten. Sauria 4 (3): 5-12 - get paper here
  • Manthey, U. 1982. Die Gattung Ptychozoon, Teil 1 - Mit einem Bestimmungsschlüssel für die fünf Arten. Sauria 4 (2): 11-17 - get paper here
  • Manthey, U. & Denzer, W. 1982. Exkursion am Mt. Kinabalu (4101 m), Nordborneo, Teil 2: Herpetologische Eindrücke. Herpetofauna 4 (21): 11-19 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Min, Pui Yong and Indraneil Das. 2012. A significant range extension for the Kinabalu Parachute Gecko, Ptychozoon rhacophorus (Boulenger, 1899) (Squamata: Gekkonidae) and a new state record from Sarawak, northwestern Borneo. Herpetology Notes 6: 177-179 - get paper here
  • Rafe M. Brown, Cameron D. Siler, Indraneil Das, Yong Min 2012. Testing the phylogenetic affinities of Southeast Asia’s rarest geckos: Flap-legged geckos (Luperosaurus), Flying geckos (Ptychozoon) and their relationship to the pan-Asian genus Gekko. Molecular Phylogenetics and Evolution 63: 915-921 - get paper here
  • Rösler, H. 2018. Haftbar -- Die einzigartige Welt der Geckos. Begleitheft zur Ausstellung im Naturkundemuseum Erfurt Mai-Aug 2018, 96 pp.
  • SUMONTHA, MONTRI; OLIVIER S.G. PAUWELS, KIRATI KUNYA, CHAIWAT LIMLIKHITAKSORN, SIRICHAI RUKSUE, APIRAT TAOKRATOK, MICHEL ANSERMET & LAWAN CHANHOME 2012. A new species of Parachute Gecko (Squamata: Gekkonidae: genus Ptychozoon) from Kaeng Krachan National Park, western Thailand. Zootaxa 3513: 68–78 - get paper here
  • Wood Jr, Perry L.; Xianguang Guo, Scott L. Travers, Yong-Chao Su, Karen V. Olson, Aaron M. Bauer, L. Lee Grismer, Cameron D. Siler, Robert G. Moyle, Michael J. Andersen, Rafe M. Brown 2019. Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. bioRxiv 717520 - get paper here
 
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