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Gekko sorok (DAS, LAKIM, KANDAUNG, 2008)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymLuperosaurus sorok DAS, LAKIM, KANDAUNG 2008
Gekko (Rhacogekko) sorok — WOOD et al. 2019 (incertae sedis)
Gekko (Rhacogekko) sorok — WOOD et al. 2020
Luperosaurus sorok — FUKUYAMA et al. 2022 
DistributionMalaysia (Borneo: Crocker Range Park, Sabah; Lambir Hills National Park)

Type locality: near bank of Sungai Bariawa, Crocker Range Park (05°24'177"N, 116°07'897"E), Keningau District, Sabah, Malaysia (Borneo).  
Reproductionoviparous 
TypesHolotype: SP 06618, female, collected 19 February 2002 by Pius Kandaung. 
DiagnosisDiagnosis: A small species of Luperosaurus (SVL 34.7 mm), diagnosable from congeneric species by the possession of the following combination of characters: body robust; head narrower than body; auricular open- ing oval-squarish; subrictal tubercles present; two intersupranasals contacting rostral; rostral contacts nares; nine supralabials to the middle of the orbital; 14/15 supralabials to posterior edge of orbit; 11 infralabials; 1–3 anteriormost chin scales subequal, larger than those to their posterior; dorsal body scales rounded, convex and granular; ca. 104 dorsal scale rows; 45 ventral scale rows; ca. 149 midbody scale rows; ventrolateral body tubercles spinose; three distal scansors deeply notched; tail fringe with distinct serrations; and pale rounded spots present on middorsum [from DAS et al. 2008]

Diagnosis. The species can be diagnosed from other species of Luperosaurus by a combination of the following characteristics: small body (SVL 28.7–34.5 mm); subrictal tubercles present; rostral contacts nares; dorsal body scales rounded, convex, and granular; ventrolateral body tubercles spinose; three distal scansors deeply notched; tail fringe with distinct serrations; 14 or 15 supralabials; 11–13 infralabials; 8–10 scansors on toe IV; 149–174 midbody scales; 45–47 precloacofemoral pores in males; and pale rounded spots present on middorsum. (Fukuyama et al. 2022)

Coloration in Life (from all referred specimens). Dorsal ground color of head, body, limbs, and tail reddish brown to light brown; many pale, irregularly shaped markings of various sizes on head, body, limbs, and tail; four black inverse V-shaped markings between limb insertions; enlarged and spinose body scales light orange or light brown. Ventral ground color of head, body, limbs, and tail light brown with scattered dark irregularly shaped small markings; palm and sole orange. Roof of the mouth dark blue; throat black; yellow-orange jaws and red tongue (Fig. 3D). (Fukuyama et al. 2022)

Intraspecific Variation.—Our specimens and the holotype are generally similar in morphology (Table 3). The two male specimens (SRC 00952 and 00954) were smaller and slenderer than the two female specimens (SRC 00872 and the holotype: SP 06618; 28.7 and 31.8 mm vs. 34.5 and 34.7 mm in SVL) and had relatively longer tails (TL 82 and 74% of SVL vs. 72 and 69% of SVL) and a higher number of midbody scales (172 and 174 vs. 150 and 149) and lower number of internasals contacting rostral (1 vs. 2). Also, only males have precloacofemoral pores (45 and 47). (Fukuyama et al. 2022) 
CommentLuperosaurus sorok was preliminarily assigned to the new subgenus Rhacogekko within Gekko. See Wood et al. 2019 for details. 
EtymologyRhacogekko is derived from the Greek noun rhakos, meaning "rag" or "wrinkle" in relation to the rounded lobe-like fringes or wrinkles on the lateral folds of the body.

The species name is derived from Malay, meaning "to hide", in reference to its presumed ecologically cryptic nature. 
References
  • Das, Indraneil; Lakim, Maklarin; Kandaung, Pius. 2008. New species of Luperosaurus (Squamata: Gekkonidae) from the Crocker Range Park, Sabah, Malaysia (Borneo). Zootaxa 1719: 53-60 - get paper here
  • Fukuyama, Ibuki; Mohamad Yazid Hossman, Kanto Nishikawa 2022. Rediscovery of Luperosaurus sorok Das, Lakim, and Kandaung, 2008 (Squamata, Gekkonidae) With Notes on Its Taxonomy and Natural History. Journal of Herpetology 56 (2): 241–248 - get paper here
  • Wood Jr, Perry L.; Xianguang Guo, Scott L. Travers, Yong-Chao Su, Karen V. Olson, Aaron M. Bauer, L. Lee Grismer, Cameron D. Siler, Robert G. Moyle, Michael J. Andersen, Rafe M. Brown 2019. Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. bioRxiv 717520 [preprint] - get paper here
  • Wood, P. L., Guo, X., Travers, S. L., Su, Y. C., Olson, K. V., Bauer, A. M., Grismer, L. L., Siler, C. D., Moyle, R. G., Andersen, M. J. and Brown, R. M. 2020. Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. Molecular Phylogenetics and Evolution 146: 106731 [corrigendum: MPE 164: 107255] - get paper here
 
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