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Geophis turbidus PAVÓN-VÁZQUEZ, CANSECO-MÁRQUEZ & NIETO-MONTES DE OCA, 2013

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymGeophis turbidus PAVÓN-VÁZQUEZ, CANSECO-MÁRQUEZ & NIETO-MONTES DE OCA 2013
Geophis rostralis — WEBB & FUGLER 1957
Geophis cf. rostralis — BOGERT & PORTER 1966
Geophis carinosus — DOWNS 1967
Geophis dubius — PÉREZ-HIGAREDA & SMITH 1988 
DistributionMexico (N Puebla)

Type locality: 3.5 km W of Xocoyolo, municipality of Cuetzalan, Puebla, Mexico (19859.4320N, 97833.3250W; datum 1⁄4 WGS84), 1225 m elevation.  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: MZFC 27254 (field number IFP 105), female, collected 24 March 1998 by Israel Fentanes Patiño, (Fig. 2).
Paratypes.—Ten specimens, all from the Sierra Norte region of Puebla, Mexico: four from the municipality of Cuetzalan (one [EBUAP 1021] from 2 km NW of Xocoyolo, near Vista Hermosa [19858.6770N, 97832.2940W], 1345 m elevation; one [EBUAP 521] from 5.5 km SSW of Cuetzalan; one [MZFC 27253] from hillside in Xocoyolo, on road to Apulco River [19857057.600N, 97832013.100W]; and one [EBUAP 1022] from La Loma del Chivo, on the road to Tacopixacta [19858.4100N, 97830.9860W], 1215 m elevation); four from the municipality of Tepango de Rodr ́ıguez (three [CNAR 6886– 6888] from Tepango de Rodr ́ıguez, and one [CNAR 6889] from 5 km S of Tepango de Rodr ́ıguez); one (KU 39642) from 16.09 km SW of Villa Juárez (Xicotepec de Juárez), and one (CNAR 8233) from 3 km N of Zacapoaxtla, on road to Cuetzalan. 
DiagnosisDiagnosis (short): G. turbidus possesses most of the diagnostic characters of the G. dubius group but differs from all other species of Geophis by having the loreal shorter than the combined prenasal and postnasal length, one postocular scale, one supraocular scale, the fifth supralabial and parietal in contact, dorsal scales in 17 rows, dorsal scales smooth throughout the length of the body or, if keeled, not anteriorly to the posterior fourth of the body, 129–140 ventrals in females and 125–139 in males, 26–31 subcaudals in females and 34–39 in males, 159–170 ventrals plus subcaudals in females and 161–174 in males, 9 maxillary teeth, anterior tip of the maxilla toothless, an overall dark dorsum, a predominantly cream venter, and a short tail in males.


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CommentSynonymy: after PAVÓN-VÁZQUEZ et al. 2013.

The status of the species is still uncertain 
EtymologyThe specific name comes from the Latin word turba, meaning confusion, turmoil, and makes reference to the past confusion regarding the taxonomic status of the new species. 
References
  • Canseco-Márquez L, Pavón-Vázquez CJ, López-Luna MA, Nieto-Montes de Oca A 2016. A new species of earth snake (Dipsadidae, Geophis) from Mexico. ZooKeys 610: 131-145. doi: 10.3897/zookeys.610.8605 - get paper here
  • Heimes, P. 2016. Snakes of Mexico. Chimaira, Frankfurt, 572 pp
  • Pavón-Vázquez, Carlos J.; Luis Canseco-Márquez, and Adrián Nieto-Montes de Oca 2013. A New Species in the Geophis dubius Group (Squamata: Colubridae) from Northern Puebla, México. Herpetologica 69 (3): 358-370. - get paper here
  • Woolrich-Piña, G. A., E. García-Padilla, D. L. DeSantis, J. D. Johnson, V. Mata-Silva, and L. D. Wilson 2017. The herpetofauna of Puebla, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 4(4): 791–884 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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