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Geophis annuliferus (BOULENGER, 1894)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Western Snail-Eating Snake
S: Caracolera Occidental 
SynonymTropidodipsas annulifera BOULENGER 1894
Geophis tecpanecus DUGES 1896
Atractus tecpanicus [sic] — BOULENGER 1897 (in Zoological Record)
Geatractus tecpanecus — DUGES 1898: 52
Tantilla nelsoni SLEVIN 1926
Tropidodipsas malacodryas SHANNON & HUMPHREY 1959
Tropidodipsas annulifera — SCOTT 1967
Sibon annuliferus — KOFRON 1985
Tantilla nelsoni SLEVIN 1926: 200
Exelencophis nelsoni — SMITH 1942: 33
Tropidodipsas occidentalis — CAMPBELL & SIMMONS 1962: 197
Sibon annuliferus — LINER 1994
Tropidodipsas annulifera — LINER 2007
Tropidodipsas annulifera — FLORES-VILELA et al. 2010
Sibon annulifera — PYRON & BURBRINK 2013
Tropidodipsas annulifera — WALLACH et al. 2014: 746
Tropidodipsas annuliferus — FELDMAN et al. 2015
Tropidodipsas annulifera — WILLIAMS & ARCE 2021
Geophis annuliferus — GRÜNWALD et al. 2021 
DistributionMexico (Sinaloa, Nayarit, Jalisco, Colima, Michoacan, Guerrero)  
Reproductionoviparous 
TypesHolotype: BMNH 1946.1.6.4
Holotype: CAS 58680 [nelsoni]
Holotype: IUMNH 75787 (FAS 12487) Mexico: 4-8 mi E San Blas, Nayarit; F. A. Shannon and F. L. Humphrey; July 18, 1957. [malacodryas]
Holotype: MDUG HE 1139 (a skull, seven dorsal vertebra and three ribs [tecpanecus] 
Diagnosis 
CommentSynonymy after KOFRON 1985. AMARAL (1929) considered Tropidodipsas annulifera as a synonym of Tropidodipsas (= Sibon) anthracops which he renamed Sibynomorphus anthracops. 
EtymologyNamed after the diminutive form (“annul-”) of Latin “anus” = ring and the verb “fero, ferre” = carry. 
References
  • Amaral,A. do 1930. Estudos sobre ophidios neotropicos XVIII. Lista remissiva dos ophidios da região neotropica. Mem. Inst. Butantan 4: 126-271 [1929] - get paper here
  • Boulenger, George A. 1894. Catalogue of the snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ. British Mus. (Nat. Hist.), London, xi, 382 pp. - get paper here
  • Campbell, H. W. & Simmons, R.S. 1962. Notes on some reptiles and amphibians from western Mexico. Bull. S. Calif. Acad. Sci. 61 (4): 193-203
  • Dugès, ALFREDO 1896. Geophis tecpanecus. Naturaleza 2nd Ser. 2: 455-456 - get paper here
  • Dugès, A. 1898. Un nuevo género de ofidio. La Naturaleza, (2)3: 52 - get paper here
  • FLORES-VILLELA, OSCAR; CÉSAR A. RÍOS-MUÑOZ, GLORIA E. MAGAÑA-COTA & NÉSTOR L. QUEZADAS-TAPIA 2016. Alfredo Dugès’ type specimens of amphibians and reptiles revisited. Zootaxa 4092 (1): 033–054 - get paper here
  • FLORES-VILLELA, OSCAR; VICTOR HUGO REYNOSO & GLORIA MAGAÑA-COTA 2010. More notes on Alfredo Dugès’ names for Mexican reptilian taxa. [Chalinocnemis, Spasmocnemis, Sceloporus westphalii nomina nuda]. Zootaxa 2600: 66–68 - get paper here
  • Goldberg, S. R., and C. R. Bursey. 2017. Tropidodipsas annulifera Boulenger, 1984. Parasite. Mesoamerican Herpetology 4(2): 441 - get paper here
  • Grünwald CI, Toribio-Jiménez S, Montaño-Ruvalcaba C, Franz-Chávez H, Peñaloza-Montaño MA, Barrera-Nava EY, Jones JM, Rodriguez CM, Hughes IM, Strickland JL, Reyes-Velasco J 2021. Two new species of snail-eating snakes of the genus Tropidodipsas (Serpentes, Dipsadidae) from southern Mexico, with notes on related species. Herpetozoa 34: 233-257 - get paper here
  • Grünwald, Christoph Imre, Nadia Pérez-Rivera, Iván Trinidad Ahumada-Carrillo, Héctor Franz-Chávez and Brandon Thomas La Forest. 2016. New distributional records for the herpetofauna of Mexico. Herpetological Review 47 (1): 85-90 - get paper here
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Hardy, L.M., & McDiarmid, R.W. 1969. The amphibians and reptiles of Sinaloa, Mexico. Univ. Kansas Publ. Mus. Nat. Hist. 18 (3): 39-252. - get paper here
  • Heimes, P. 2016. Snakes of Mexico. Chimaira, Frankfurt, 572 pp
  • Kofron, C.P. 1988. Systematics of neotropical gastropod-eating snakes: the sartorii group of the genus Sibon. Amphibia-Reptilia 9: 145-168 - get paper here
  • Lemos-Espinal JA, Smith GR 2020. A checklist of the amphibians and reptiles of Sinaloa, Mexico with a conservation status summary and comparisons with neighboring states. ZooKeys 931: 85-114 - get paper here
  • Lemos-Espinal JA, Smith GR, Pierce LJS, Painter CW 2020. The amphibians and reptiles of Colima, Mexico, with a summary of their conservation status. ZooKeys 927: 99-125 - get paper here
  • Liner, Ernest A. 2007. A CHECKLIST OF THE AMPHIBIANS AND REPTILES OF MEXICO. Louisiana State University Occasional Papers of the Museum of Natural Science 80: 1-60 - get paper here
  • Loc-barragán J.A., D., Lazcano y G.A Woolrich-Piña. 2018. Notes on the Herpetofauna of Nayarit, Mexico 2: Amphibians and Reptiles of the Municipality of Compostela. Bulletin of the Chicago Herpetological Society 53(10): 205-212
  • Loc-Barragán JA, Smith GR, Woolrich-Piña GA, Lemos-Espinal JA 2024. An updated checklist of the amphibians and reptiles of Nayarit, Mexico with conservation status and comparison with adjoining States. Herpetozoa 37: 25-42 - get paper here
  • Loc-Barragán, Jesús Alberto and David Lazcano 2018. Notes on the Herpetofauna of Nayarit, Mexico 1: Amphibians and Reptiles of the Municipality of Tecuala. Bulletin of the Chicago Herpetological Society 53(4):73-80
  • Palacios-Aguilar, Ricardo & OSCAR FLORES-VILLELA 2018. An updated checklist of the herpetofauna from Guerrero, Mexico. Zootaxa 4422 (1): 1-24 - get paper here
  • Pyron, R. Alexander; Frank T. Burbrink 2013. Early origin of viviparity and multiple reversions to oviparity in squamate reptiles. Ecology Letters 17 (1): 13–21 (published online 2013, in print 2014), DOI: 10.1111/ele.12168 - get paper here
  • Scott Jr., N.J. 1967. The colubrid snake, Tropidodipsas annulifera, with reference to the status of Geatractus, Exelencophis, Chersodromus annulatus, and Tropidodipsas malacodryas. Copeia 1967 (2): 280-287 - get paper here
  • Shannon,F.A. & Humpfrey,F.L. 1959. Two new species of Tropidodipsas from the west coast of Mexico. Herpetologica 15 (4): 217-222 - get paper here
  • Slevin, Joseph Richard 1926. Expedition to the Revillagigedo Islands, Mexico, in 1925, III. Notes on a collection of reptiles and amphibians from the Tres Marias and Revillagigedo Islands, and the West Coast of Mexico, with description of a new species of Tantilla. Proc. Cal. Acad. Sci. 15 (3): 195-207 - get paper here
  • Smith, H.M. 1942. A resume of Mexican snakes of the genus Tantilla. Zoologica 27: 33-42. - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Webb, R.G. 1984. Herpetogeography in the Mazatlán-Durango Region of the Sierra Madre Occidental, Mexico. Vetrebrate Ecology and Systematics - A ribute to Henry S. Fitch; Museum of Natural History, University of Kansas, Lawrence, pp. 217-241
  • WILLIAMS, SPENCER & JOSE ANGEL ARCE 2021. New Distributional Records from Mixtlán, an Undersampled Municipality in Jalisco, Mexico. Herpetological Review 52 (2): 359–361
  • Woolrich-Piña, Guillermo A, Paulino Ponce-Campos, Jesús Loc-Barragán, Juan Pablo Ramírez-Silva, Vicente Mata-Silva, Jerry D. Johnson, Elí García-Padilla and Larry David Wilson. 2016. The herpetofauna of Nayarit, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 3 (2): 376-448 - get paper here
 
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