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Geophis sanniolus (COPE, 1866)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
SubspeciesGeophis sanniolus neilli (HENDERSON, HOEVERS & WILSON 1977)
Geophis sanniolus sanniolus (COPE 1866) 
Common NamesE: Pigmy Snail Sucker
S: Caracolera Pigmea 
SynonymMesopeltis sanniolus COPE 1866: 318
Leptognathus sanniola — BOCOURT 1908
Sibynomorphus sanniola — SCHMIDT & ANDREWS 1936
Sibynomorphus sanniolus — GAIGE 1936
Sibon neilli HENDERSON, HOEVERS & WILSON 1977
Sibon sanniola — LEE 2000: 329
Sibon sanniolus — LINER 1994
Sibon sanniola — CAMPBELL 1998
Sibon sanniolus — MCCRANIE 2006
Sibon sanniolus — WALLACH et al. 2014: 669
Sibon sanniola — HEIMES 2016: 309
Geophis sanniolus — GRÜNWALD et al. 2021

Sibon sanniolus neilli HENDERSON, HOEVERS & WILSON 1977
Sibon neilli HENDERSON, HOEVERS & WILSON 1977
Sibon sanniola neilli — LEE 2000: 330
Sibon sanniola neilli — STAFFORD 2000: 239 
DistributionMexico (Yucatan, Campeche, Quintana Roo), Belize, Guatemala

Type locality: Yucatan

neilli: Belize; Type locality: vicinity of Belize City, Belize District, Belize,  
Reproductionoviparous 
TypesHolotype: USNM 6564, male, from Yucatan.
Holotype: MPM 8929 (Milwaukee Public Museum), a male, collected July 1974 by Ernest Smith. [neilli] 
DiagnosisDiagnosis: S. neilli differs from all other Sibon, except S. sanniola, in not having the loreal enter the orbit and in the high number of supralabials. It differs from male S.sanniola in having a slightly higher number of ventrals and subcaudals and is strikingly different in color pattern. S. neilli has 34 dark brown body bands whereas S. sanniola has a middorsal row of spots. In addition, S. neilli has the 7th supralabial in contact with the lower postocular and primary temporal, but not in contact with the secondary temporal. This condition occurs occasionally in S. sanniola, but in no other species of Sibon. 
CommentSynonymy after VILLA et al. 1988 (but see KOFRON 1990). 
EtymologyNamed after Latin sannio, one who mimics in mockery, a harlequin, buffoon + Latin -olus, suffix dimminutive. [?].

Geophis sanniolus neilli was named after Wilfred Trammell Neill Jr. (1922–2001), US American herpetologist. 
References
  • Aguilar-López JL, Luría-Manzano R, Pineda E, Canseco-Márquez L 2021. Selva Zoque, Mexico: an important Mesoamerican tropical region for reptile species diversity and conservation. ZooKeys 1054: 127-153 - get paper here
  • Barbour, T., and L. J. Cole. 1906. Vertebrata from Yucatan. Reptilia, Amphibia and Pisces. Bull. Mus. Comp. Zool. Harvard 50: 146-159 - get paper here
  • Calderon, R.; Cedeño-Vázquez, J.R. & Pozo, C. 2003. New distributional records for amphibians and reptiles from Campeche, Mexico. Herpetological Review 34 (3): 269-272 - get paper here
  • Campbell, J.A. 1998. Amphibians and reptiles of northern Guatemala, the Yucatán, and Belize. Norman: University of Oklahoma Press, xiii + 380 pp. - get paper here
  • Cope, E.D. 1867. Fifth contribution lo the herpetology of tropical America. Proc. Acad. nat. Sci. Philadelphia, 18 [1866]: 317-323 - get paper here
  • Gaige,H. 1936. Some reptiles and amphibians from Yucatan and Campeche, Mexico. Carnegie Inst. Wash. Publ., (457): 289-304.
  • Grünwald CI, Toribio-Jiménez S, Montaño-Ruvalcaba C, Franz-Chávez H, Peñaloza-Montaño MA, Barrera-Nava EY, Jones JM, Rodriguez CM, Hughes IM, Strickland JL, Reyes-Velasco J 2021. Two new species of snail-eating snakes of the genus Tropidodipsas (Serpentes, Dipsadidae) from southern Mexico, with notes on related species. Herpetozoa 34: 233-257 - get paper here
  • Harvey, Michael B.; Gilson Rivas Fuenmayor, José Rances Caicedo-Portilla, and José Vicente Rueda-Almonacid 2009. Systematics of the Enigmatic Dipsadine Snake Tropidodipsas perijanensis Alemán (Serpentes: Colubridae) and Review of Morphological Characters of Dipsadini. Herpetological Monographs 22 (1): 106-132 - get paper here
  • Heimes, P. 2016. Snakes of Mexico. Chimaira, Frankfurt, 572 pp
  • Henderson, R. W., HOEVERS, L. G., & WILSON, L. D. 1977. A new species of Sibon (Reptilia, Serpentes, Colubridae) from Belize, Central America. Journal of Herpetology 11 (1): 77-79. - get paper here
  • Kofron, C.P. 1990. Systematics of Neotropical gastropod-eating snakes: the dimidiata group of the genus Sibon, with comments on the nebulata group. Amphibia-Reptilia 11: 207-223 - get paper here
  • Köhler, G. 2008. Reptiles of Central America. 2nd Ed. Herpeton-Verlag, 400 pp.
  • Köhler, Gunther, J. Rogelio Cedeño-Vázquez, Till Kirstein and Pablo Beutelspacher-García. 2016. The Chetumal Snake Census: generating biological data from road-killed snakes. Part 2. Dipsas brevifacies, Sibon sanniolus, and Tropidodipsas sartorii. Mesoamerican Herpetology 3 (3): 689–705 - get paper here
  • Lee, J. C. 2000. A field guide to the amphibians and reptiles of the Maya world. Cornell University Press, Ithaca,
  • Lee, J.C. 1996. The amphibians and reptiles of the Yucatán Peninsula. Comstock, Cornell University Press, Ithaca, 500 pp.
  • McCOY, C. J. 1986. Results of the Carnegie Museum of Natural History Expeditions to Belize. I. Systematic status and geographic distribution of Sibon neilli (Reptilia, Serpentes). Annals of the Carnegie Museum 55: 117-123. - get paper here
  • McCOY, C. J., CENSKY, E. J., & VAN DE VENDER, R. R. 1986. Distribution records for amphibians and reptiles in Belize, Central America. Herpetological Review 17:28-29. - get paper here
  • McCranie, J.R. 2007. A second new species of Sibon (Squamata: Colubridae) from La Mosquitia, Northeastern Honduras. Herpetologica 63 (2): 213-218 - get paper here
  • McCranie, James R. 2006. New species of Sibon (Squamata: Colubridae) from Northeastern Honduras. Journal of Herpetology 40 (1): 16-21 - get paper here
  • Pérez-Higareda, Gonzalo, Marco A. López-Luna, and Hobart M. Smith 2002. A new snake related to Sibon sanniola (Serpentes: Dipsadidae) from Los Tuxtlas, Veracruz, Mexico. Bull. Maryland Herp. Soc. 38(2):62-65 - get paper here
  • Peters , J. A. 1960. The snakes of the subfamily Dipsadinae. Misc. Publ. Mus. Zool., Univ. Michigan (114): 224 pp. - get paper here
  • Pizzatto, Lígia; Maurício Cantor, Juliana Lima de Oliveira, Otavio A.V. Marques, Vinicius Capovilla, and Marcio Martins 2008. Reproductive Ecology of Dipsadine Snakes, With Emphasis on South American Species. Herpetologica 64 (2): 168-179 - get paper here
  • ROVITO, SEAN M.; THEODORE J. PAPENFUSS 2012. A new species of Sibon (Squamata: Colubridae) from the mountains of eastern Guatemala. Zootaxa 3266: 62–68 - get paper here
  • Schmidt, K. P, & ANDREWS, E. W. 1936. Notes on snakes from Yucatán. Field Mus. Nat Hist. Zool. Ser. 20: 167-187. - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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