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Gloydius cognatus (GLOYD, 1977)

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Higher TaxaViperidae, Crotalinae, Colubroidea, Alethinophidia, Serpentes, Squamata (snakes) 
Common NamesE: Alashan pitviper
G: Alashan-Otter
Chinese: 阿拉善蝮 
SynonymAgkistrodon halys cognatus GLOYD 1977
Agkistrodon halys cognatus — WELCH 1994: 11
Gloydius halys cognatus — GUMPRECHT et al. 2004
Gloydius halys cognatus — WAGNER et al. 2015
Gloydius cognatus — SHI et al. 2016
Gloydius cognatus — SHI et al. 2018
Gloydius cognatus — ASADI et al. 2019 
DistributionNC China (Gansu)

Type locality: Choni, on the Tao River, Gansu Province, China.  
TypesHolotype: USNM 68586. Paratypes. USMN 68504, 68506, 68541 
DiagnosisDiagnosis. A small viper of the genus, up to 590 mm total length. Supralabial scales 7-8. Dorsal scales in 23 (rarely 21) rows around mid-body; ventral scales between 153-165; subcaudal scales paired, between 36-54. Body with 29- 43 dark transverse bands, each 4-5 scales wide and extending down to scale row 4 to 2 [WAGNER et al. 2015].

Diagnosis. Total length up to 590 mm. Supralabials 7 – 8; dorsal scale in 23 rows at midbody, sometimes 21; ventrals 153 – 165; subcaudals 36 – 54 [ORLOV & BARABANOV 1999]. 

Synonymy: Zhao 2006 listed Gloydius halys cognatus, liupanensis, qinlingensis, and stejnegeri as synonyms of either G. strauchi or G. intermedius.

Activity Pattern. Diurnal and night. 
  • Asadi, Atefeh; Claudine Montgelard, Masoud Nazarizadeh, Akram Moghaddasi,<br />Faezeh Fatemizadeh, evgeniy simonov, Haji Gholi Kami & Mohammad Kaboli 2019. Evolutionary history and postglacial colonization of an Asian pit viper (Gloydius halys caucasicus) into transcaucasia revealed by phylogenetic and phylogeographic analyses. Scientific Reports 9:1224 - get paper here
  • Gloyd, H.K. 1977. Descriptions of new taxa of crotalid snakes from China and Ceylon (Sri Lanka). Proc. Biol. Soc. Washington 90: 1002-1015 - get paper here
  • Gumprecht, A.; Tillack, F.; Orlov, N.L.; Captain, A. & Ryabow, S. 2004. Asian pitvipers. Geitje Books, Berlin, 368 pp.
  • Shi, J., Yang, D., Zhang, W. & Ding, L. 2016. Distribution and infraspecies taxonomy of Gloydius halys-Gloydius intermedius complex in China (Serpentes: Crotalinae). Chinese J. Zool. 51: 777–798
  • Shi, Jingsong; Diancheng Yang, Wuyuan Zhang, Lifang Peng, Nikolai L. Orlov, Fan Jiang, Li Ding, Mian Hou, Xinlei Huang, Song Huang, Pipeng Li 2018. A New Species of the Gloydius strauchi Complex (Crotalinae: Viperidae: Serpentes) from Qinghai, Sichuan, and Gansu, China. Russ. J. Herpetol. 25 (2): 126-138 - get paper here
  • Wagner, Philipp; Tiutenko, Arthur; Mazepa, Glib; Borkin, Leo J.; Simonov, Evgeniy 2015. Alai! Alai! – a new species of the Gloydius halys (Pallas, 1776) complex (Viperidae, Crotalinae), including a brief review of the complex. Amphibia-Reptilia 37 (published 31 Dec 2015) DOI: 10.1163/15685381-00003026 - get paper here
  • Welch, K. R. G. 1994. Snakes of the World. A Checklist. I. Venomous snakes. KCM Books, Somerset, England.
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