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Goggia incognita HEINICKE, TURK & BAUER`, 2017

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common Names 
SynonymGoggia incognita HEINICKE, TURK & BAUER 2017
Diplodactylus lineatus GRAY 1845 (part)
Phyllodactylus lineatus — SMITH 1849 (part)
Phyllodactylus lineatus lineatus — HEWITT 1937 (part)
Goggia lineata — BAUER, GOOD, and BRANCH 1997 (part) 
DistributionRepublic of South Africa (Western Cape)

Type locality: South Africa, Western Cape, Jacobsbaai (32° 59' 19" S, 17° 52' 37" E).  
Reproduction 
TypesHolotype: CAS 224024: adult male. Collected by A. M. Bauer, R. A. Sadlier, A. Whiting, 9 September 2001.
Paratypes. CAS 224022 (adult female), CAS224023 (adult female): same data as holotype. CAS 176047 (adult female): Same locality as holotype. Collected by A. M. Bauer, 23 May 1990. CAS 206692 (adult male), CAS 206697 (adult female): Same locality as holotype. Collected by A. M. Bauer, A. C. Lamb, J. L. Wright, P. Moler, R. D. Babb, 11 July 1998. MCZ R-192442 (adult female): South Africa, Western Cape, Mauritzbaai (32° 58' 39.8" S, 17° 52' 51.9" E). Collected by M. P. Heinicke, A. M. Bauer, T. Gamble, D. Zarkower, J. Marais, A. Kuhn, E. Frietas, R. Skinner, 2 August 2013. MCZ R 192395 (adult females), MCZ R-194417 (adult male): South Africa, Western Cape, 1 km S Jacobsbaai (32° 59 ́ 19.1 ́ ́ S 17° 52 ́ 36 ́ ́ E). Collected by M. P. Heinicke, A. M. Bauer, T. Gamble, D. Zarkower, J. Marais, A. Kuhn, E. Frietas, R. Skinner, 24 July 2013. MCZ R-192446 (subadult male), MCZ R-194450 (adult male): South Africa, Western Cape, 4.3 km S Jacobsbaai (33° 0 ́ 20.9 ́ ́ S, 17° 52 ́ 46.5 ́ ́ E). Collected by M. P. Heinicke, A. M. Bauer, T. Gamble, D. Zarkower, J. Marais, A. Kuhn, E. Frietas, R. Skinner, 2 August 2013. 
CommentSimilar species: Goggia lineata, as which it has been identified in the older literature.

Distribution: The distribution of this species comprises what was formerly considered the southern half of the distribution of G. lineata. The boundary between the two species appears to be at the Knersvlakte, a quartz plain lying between the Cape Fynbos and Little Namaqualand ecoregions, near the border between the Western Cape and Northern Cape provinces, with G. incognita sp. nov. occurring only to the south in the Cape (Fig. 2).

Habitat: terrestrial, nocturnal species that shelters by day under a wide range of available debris, including under stones, in plant litter, and under or within dead Aloe stems (Branch & Bauer 1995, Branch & Braack 1989, Branch 1998). At the type locality, along the coast, individuals may be encountered both under boulders and under strand debris. At Farm Buffelskloof, in the Little Karoo, no individuals were found sheltering under the local conglomerate rock, but one individual was found inside a dead Aloe.

Sympatry: Afrogecko porphyreus, Pachydactylus geitje (near Jacobsbaai); Goggia hewitti and Pachydactylus geitje at Farm Buffleskloof. 
EtymologyThe specific epithet is from the Latin word incognitus, meaning “not known”. The English phrase “going incognito” refers to remaining hidden or disguised. The name is chosen to reflect the 150+ year time period in which this species has remained hidden within what were considered nominotypical populations of Goggia lineata. It additionally reflects the natural history of the species, as members of the species are typically inconspicuous and hidden under cover objects by day. The name is used as an adjective. 
References
  • Bauer A M. Good D A. Branch W R. 1997. The taxonomy of the Southern African leaf-toed geckos (Squamata: Gekkonidae), with a review of Old World "Phyllodactylus" and the description of five new genera. Proc. Cal. Acad. Sci. 49 (14): 447-497. - get paper here
  • Gray, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp. - get paper here
  • HEINICKE, MATTHEW P.; DILARA TURK, AARON M. BAUER 2017. Molecular phylogeny reveals strong biogeographic signal and two new species in a Cape Biodiversity Hotspot endemic mini-radiation, the pygmy geckos (Gekkonidae: Goggia) Zootaxa 4312 (3): 449–470 - get paper here
  • Hewitt, J. 1937. Descriptions of South African lizards. Ann. natal. Mus. 8: 199-209
  • Smith, A. 1849. Illustrations of the Zoology of South Africa. 3 (Reptiles). Smith, Elder, and Co., London [facsimile published by Winchester Press, Johannesburg, 1977] - get paper here
 
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