Goniurosaurus toyamai GRISMER, OTA & TANAKA, 1994

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Higher Taxa	Eublepharidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies
Common Names	E: Iheyajima Leopard Gecko Japanese: Iheya-Tokagemodoki
Synonym	Goniurosaurus kuroiwae toyamai GRISMER, OTA & TANAKA 1994: 328 Goniurosaurus kuroiwae orientalis — OTA 1989: 230 (part.) Goniurosaurus kuroiwae toyamai GRISMER et al. 1994 Goniurosaurus toyamai — DICKHOFF 2004 Goniurosaurus toyamai — SEUFER et al. 2005 Goniurosaurus toyamai — ZIEGLER et al. 2008 Goniurosaurus toyamai — KURITA et al. 2017 Goniurosaurus kuroiwae toyamai — KURITA & TODA 2024 Goniurosaurus toyamai — KURITA & TODA 2024
Distribution	Japan (Iheyajima) Type locality: Iheyajima island, Okinawa, Japan
Reproduction	oviparous
Types	Holotype: KUZ 9983
Diagnosis	Diagnosis: Goniurosaurus kuroiwae toyamai differs from all other subspecies of G. kuroiwae in its overall robust body stature and greater mean number of paravertebral tubercles (37.2:34-42). It differs further from G. k. yamashinae, G. k. orientalis, and G. k. kuroiwae in that adults lack interspace mottling; from G. k. orientalis and G. k. kuroiwae in lacking lineate tendencies in its dorsal pattern and having a lower mean number of eyelid fringe scales (55.6:53-59); from G. k. orientalis by having a slightly higher mean number of postmental scales (4.6:3-5); from G. k. kuroiwae in having a complete dorsal banding pattern; from G. k. yamashinae by having enlarged scales at the base of its digits, orange-pink juvenile color pattern, blood-red iris, a higher mean number of supralabial scales (9.3:8-10), and a lower mean number of fourth toe lamellae (17.3:16-20); and from G. k. splendens by lacking sharply keeled dorsal tubercles, juxtaposed and sharply raised ventral scales, the lateral extension of the posteriormost body bar onto the hind limb, and by having a greater mean number of postmental scales (4.6:3-5), midbody scales (149.8:140-158), fourth toe lamellae (17.3:16-20), and a lower mean number of scales surrounding the claw on the fourth toe (5.9:5-6). (Grismer et al. 1994) Unfortunately we had to temporarily remove additional information as this was scraped by multiple AI companies who sell that data to their customers. These details, e.g. detailed descriptions or comparisons (about 7120 characters) are available for collaborators and contributors. Please contact us for details.
Comment	This species is critically endangered. Abundance: only known from the type locality (Meiri et al. 2017). Distribution: see map in Kurita & Toda 2024: 88 (Fig. 1)
References	Aizawa, Masaya 2021. Heimlich, still und schön - Okinawa-Krallengeckos. DATZ 2021 (5): 14-23 Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here Dickhoff, A. 2004. Ein Tiger im Terrarium - Haltung und Nachzucht von Goniurosaurus araneus (GRISMER, VIETS & BOYLE 1999). Draco 5 (18): 76-81 - get paper here Grismer L L. Ota H. Tanaka S. 1994. Phylogeny, classification, and biogeography of Goniurosaurus kuroiwae (Squamata: Eublepharidae) from the Ryukyu Archipelago, Japan, with description of a new subspecies. Zoological Science (TOKYO) 11 (2): 319-335 - get paper here Grismer, L. Lee 2002. Goniurosaurus: Ancient Gekkos of the Far East. Gekko 3 (1): 22-28 Janssen, Jordi & Chris R. Shepherd 2019. Trade in Endangered and Critically Endangered Japanese Herpetofauna Endemic to The Nansei Islands Warrants Increased Protection. Current Herpetology Feb 2019 Vol. 38, No. 1: 99-109 - get paper here Kurita T, Honda M, Toda M. 2017. Species delimitation and biogeography of the Ryukyu ground geckos, Goniurosaurus kuroiwae ssp. (Squamata: Eublepharidae), by use of mitochondrial and nuclear DNA analyses. J Zool Syst Evol Res. 2017;00:1–14 - get paper here KURITA, T., & TODA, M. 2024. Genetic and Morphological Studies on Goniurosaurus kuroiwae (Squamata: Eublepharidae), with a Description of a New Species from the Northern Part of Okinawajima Island, Ryukyu Archipelago, Japan. Current Herpetology, 43(1), 86-114 - get paper here Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions 24 (2): 262-273 - get paper here Ota, Hidetoshi 1989. A review of the geckos (Lacertilia: Reptilia) of the Ryukyu Archipelago and Taiwan. in: Matsui et al., eds; Current Herpetology in East Asia: Proceedings of the Second Japan-China Herpetological Symposium Kyoto, July 1988: 222-261 - get paper here Prival, D. 2002. Five days in Chiapas (or why I will never be invited to go herping in Chiapas ever again). Sonoran Herpetologist 15 (8):86-88. - get paper here Seufer, H.; Y. Kaverkin & A. Kirschner (eds.) 2005. Die Lidgeckos. Kirschner und Seufer Verlag, 238 pp. Trape, J.-F. 2023. Guide des serpents d’Afrique occidentale, centrale et d’Afrique du Nord. IRD Éditions, Marseille, 896 pp. Wang, Ying-Yong; Jian-Huan Yang, and L. Lee Grismer 2013. A New Species of Goniurosaurus (Squamata: Eublepharidae) from Libo, Guizhou Province, China. Herpetologica 69 (2): 214-226. - get paper here Wang, Ying-Yong; Jian-Huan Yang, and Rong-Feng Cui 2010. A NEW SPECIES OF GONIUROSAURUS (SQUAMATA: EUBLEPHARIDAE) FROM YINGDE, GUANGDONG PROVINCE, CHINA. Herpetologica 66 (2): 229 - get paper here Ziegler, T.; Truong, N.Q.; Schmitz, A.; Stenke, R. & Rösler, H. 2008. A new species of Goniurosaurus from Cat Ba Island, Hai Phong, northern Vietnam (Squamata: Eublepharidae). Zootaxa 1771: 16–30 - get paper here
External links	IUCN Red List National Center for Biotechnology Information Google images

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