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Gonocephalus bornensis (SCHLEGEL, 1851)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Borneo Forest Dragon 
SynonymLophyrus bornensis [sic] SCHLEGEL 1851: 6
Tiaris miotympanum GÜNTHER 1872
Gonyocephalus miotympanum — BOULENGER 1885: 287
Gonyocephalus borneensis — BOULENGER 1885: 288
Gonocephalus borneensis — SMITH 1931: 10, 31
Goniocephalus borneensis — TAYLOR 1963: 877
Goniocephalus borneensis — GRANDISON 1972: 76
Gonyocephalus miotympanum — BOULENGER 1885
Gonyocephalus miotympanum — DE ROOIJ 1915: 106
Gonyocephalus borneensis — DE ROOIJ 1915: 106
Gonocephalus miotympanum — MANTHEY 1983
Goniocephalus borneensis — BONG HEANG 1987
Gonocephalus denzeri MANTHEY 1991 (fide MANTHEY & GROSSMANN 1997)
Gonocephalus borneensis — MANTHEY & GROSSMANN 1997: 179
Gonocephalus borneensis — MANTHEY & SCHUSTER 1999: 61
Gonocephalus bornensis [sic] — MALKMUS et al. 2002: 241
Gonocephalus bornensis [sic] — DENZER & MANTHEY 2009
Gonocephalus bornensis — MANTHEY 2010 
DistributionIndonesia (Borneo, Natuna islands), Malaysia (Sabah, Sarawak)
Thailand (reported only once near the border of Surat Thani and Nakhon Si Thammarat [fide TAYLOR 1963])

Type locality: Borneo  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesSyntypes: RMNH 3043 & 3044, two females
Hiolotype: BMNH 1946.8.19.82 [miotympanum]
Holotype: ZFMK 50527 [Gonocephalus denzeri] 
Diagnosis 
CommentIllustration in BARTS & WILMS 2003.

Synonymy: Listed as a synonym of G. bellii by WERMUTH 1967. 
EtymologyNamed after its distribution on Borneo (Indonesia). 
References
  • Ahmad, Norhayati; Eddie Ahmad, Mahathir Ratag, Edward Asrul Alimin Sinon, Brandon Don, Francer Francis, Muhammad Ridzuan Mahmod, Amshari Agimin, Daicus Belabut 2019. Amphibians and Reptiles of Imbak Canyon Study Centre and Batu Timbang Camp. Journal of Tropical Biology and Conservation 16: 25–33
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Bong Heang, Kiew 1987. An annotated checklist of the herpetofauna of Ulu Endau, Johore, Malaysia. Malayan Nature Journal 41 (2-3): 413-423.
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das, I. 1993. Jour. Sarawak Mus., 44 (65): 128
  • Das, I. & Yaakob, N. 2007. Status of knowledge of the Malaysian herpetofauna. In Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. in: L. S. L. Chua, L. G. Kirton & L. G. Saw (eds.), Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. Forest Research Institute Malaysia, Kepong, pp. 31-81
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Denzer, W. & Manthey, U. 2009. Remarks on the type specimen of Gonocephalus mjobergi Smith, 1925 (Sauria: Agamidae). Bonner zoologische Beiträge 56 (4): 255–258 - get paper here
  • Fukuyama, I; Nishikawa, K; Takata, K; Shimada, T & Hamidy, A 2019. Gonocephalus bornensis (Borneo Anglehead Lizard) Reproduction. Herpetological Review 50 (3): 575-576 - get paper here
  • Grandison, A.G.C. 1972. The Gunong Benom Expedition 1967. 5. Reptiles and amphibians of Gunong Benom with a description of a new species of Macrocalamus. Bull. Br. Mus. nat. Hist. (Zool.), London, 23: 45-101. - get paper here
  • Herlambang, A. E., Riyanto, A., Munir, M., Hamidy, A., Kimura, K., Eto, K., & Mumpuni, M. 2022. AFTER 16 YEARS: AN UPDATE CHECKLIST OF HERPETOFAUNA ON THE NATUNA ISLANDS, INDONESIA. TREUBIA, 49(2), 67-84 - get paper here
  • Janzen, Peter 2022. Herpetofauna in Sarawak, East Malaysia (Borneo) – Part 2 Reptiles. Sauria 44 (3): 3-28
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Malkmus, Rudolf 2015. Die Amphibien des Nationalparks Gunung Mulu auf Borneo. Terraria-Elaphe 2015 (2): 32-43 - get paper here
  • Manthey U 1991. Gonocephalus denzeri sp. n. (Sauria: Agamidae) aus Sarawak, Borneo. Salamandra 27 (3) 1991: 152-157 - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Manthey,U. 1983. Exkursion am Mt. Kinabalu (4101 m), Nordborneo, Teil 3: Checkliste der Herpetofauna oberhalb 600 m ü. NN. Herpetofauna 5 (23): 20-31 - get paper here
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • Schlegel,H. 1851. Descriptions de plusieurs expèces nouvelles du genre Lophyrus. Bijdr. Dierk. (Amsterdam) 3 (1): 4-6 - get paper here
  • Smith, M.A. 1931. The herpetology of Mt. Kinabalu, North Borneo, 13455 ft. Bull. Raffles Mus. No. 5: 3-32. - get paper here
  • Taylor, E.H. 1963. The lizards of Thailand. Univ. Kansas Sci. Bull. 44: 687-1077. - get paper here
  • Welton, Luke J.;Cameron D. Siler, L. L. Grismer, Arvin C. Diesmos, Jack W. Sites andRafe M. Brown 2016. Archipelago-wide survey of Philippine forest dragons (Agamidae: Gonocephalus): multilocus phylogeny uncovers unprecedented levels of genetic diversity in a biodiversity hotspot. Biological Journal of the Linnean Society, DOI: 10.1111/bij.12878 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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