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Gonocephalus chamaeleontinus (LAURENTI, 1768)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards) 
Common NamesE: Chameleon Forest Dragon, Chameleon Anglehead Lizard
G: Chamäleon-Winkelkopfagame 
SynonymIguana chamaeleontina LAURENTI 1768: 47
Agama gigantea KUHL 1820 (fide BOULENGER 1885)
Agama tigrina MERREM 1820
Lophyrus tigrinus — DUMÉRIL & BIBRON 1837: 421 (fide BOULENGER 1885)
Galeotes lophyrus SCHLEGEL 1844 (fide BOULENGER 1885)
Lophyrus sumatranus SCHLEGEL 1848 (fide BARBOUR 1912)
Gonyocephalus chamaeleontinus — BOULENGER 1885: 285
Gonyocephalus sumatranus — BOULENGER 1885: 286
Gonyocephalus chamaeleontinus — DE ROOIJ 1915: 102
Gonyocephalus sumatranus — DE ROOIJ 1915: 103
Goniocephalus chamaeleontinus — HENDRICKSON 1966: 60
Gonocephalus chamaeleontinus — MANTHEY & GROSSMANN 1997: 180
Gonocephalus chamaeleontinus — COX et al. 1998: 95
Gonocephalus chamaeleontinus — MANTHEY & SCHUSTER 1999: 58
Gonocephalus chamaeleontinus — GRISMER 2011
Gonocephalus chamaeleontinus — GRISMER & QUAH 2019 
DistributionIndonesia (Mentewei I, Sumatra, Bangka, Natuna I, Java, Nias, Sipora, Siberut, Bangkaru, Tuangku, Banyak Islands, Tanahmasa),
West Malaysia, Pahang (Pulau Tioman)

Type locality: “America” (in error)  
TypesHolotype: Iconotype (Seba. 1: 157, pl. 100 fig. 2); orginal type lost (fide Manthey & Denzer 1993: 26 who announced to designate a neotype; we are not aware of any publication in which this happened though). 
CommentSynonymy mostly after WERMUTH 1967.

Type species: Lophyrus tigrinus DUMÉRIL & BIBRON 1837 is the type species of the genus Gonocephalus KAUP 1825 (fide WERMUTH 1967). Gonocephalus GÜNTHER 1854 is a genus of Teleostei (fish). Lophyrus DUMÉRIL 1805 is preoccupied by Lophyrus POLI 1791 (Mollusca) and other genera called Lophyrus (see WERMUTH 1967: 55).

Note that the name Gonyocephalus is an emendation introduced by Wagler (1830) albeit Kaup originally used Gonocephalus and later amended it to Goniocephalus. According to ICZN 51.3.1. “Parentheses are not used when the species-group name was originally combined with an incorrect spelling or an emendation of the generic name (this applies even though an unjustified emendation is an available name with its own authorship and date [Art. 33.2.3]).” 
  • Barbour, Thomas 1912. A Contribution to the Zoögeography of the East Indian Islands. Memoirs of the Museum of Comparative Zoölogy 44 (1): 1-203 - get paper here
  • Bonetti, Mathilde 2002. 100 Sauri. Mondadori (Milano), 192 pp.
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp.
  • Dring J C M 1979. Amphibians and reptiles from northern Trengganu, Malaysia, with descriptions of two new geckos: Cnemaspis and Cyrtodactylus. Bulletin of the British Museum (NATURAL HISTORY) ZOOLOGY 34 (5): 181-241 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Evers, M. 2013. Der Traum vom Urwald im Wohnzimmer. Winkelkopfagamen (Gonocephalus) im Terrarium. Reptilia (Münster) 18 (102): 50-59 - get paper here
  • Evers, Michael 2010. Winkelkopfagamen. Gonocephalus belli, C. chamaeleontinus, G. doriae abbotti. Natur und Tier Verlag, 64 pp. - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Grismer, L.L. 2011. Amphibians and reptiles of the Seribuat Archipelago. Edition Chimaira, Frankfurt, 239 pp.
  • Grossmann, W. & Tillack, F. 2004. Pulau Tioman - Perle im Südchinesischen Meer, Teil 1. Reptilia (Münster) 9 (50): 42-49 - get paper here
  • Hallermann, J. 2005. The bizarre arboreal agamids. Reptilia (GB) (42): 8-15 - get paper here
  • Hallermann, J. 2005. Mit Hörnern, Kämmen und Gleithäuten - die bizarren Baumagamen. Reptilia (Münster) 10 (51): 18-25 - get paper here
  • Hendrickson,J.R. 1966. Observations on the fauna of Pulau Tioman and Pulau Tulai. 5. The Reptiles. Bull. Nat. Mus. Singapore 34: 53-71
  • Hien,P. Grossmann,W. & Schäfer, C. 2001. Beitrag zur Kenntnis der landbewohnenden Reptilienfauna von Pulau Tioman, West-Malaysia. Sauria 23 (4): 11-28 - get paper here
  • Kopstein, F. 1938. Ein Beitrag zur Eierkunde und zur Fortpflanzung der Malaiischen Reptilien. Bull. Raffles Mus. No 14: 81-167
  • Laurenti, J. N. 1768. Specimen medicum, exhibens synopsin reptilium emendatam cum experimentis circa venena et antidota reptilium austracorum, quod authoritate et consensu. Vienna, Joan. Thomae, 217 pp. - get paper here
  • Lim, K.K.P. & Ng, H.H. 1999. The terrestrial herpetofauna of Pulau Tioman, Peninsular Malaysia. Raffles Bull. Zool., Suppl. No. 6: 131-155 - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Terralog 7b, Edition Chimaira, Frankfurt, 168 pp.
  • Manthey, S. 2011. Agamen-Tagung 2010. Iguana Rundschreiben 24 (1): 28-32
  • Manthey, U. & DENZER, W. 1993. Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae) VI. Die chamaeleontinus-Gruppe. Sauria 15 (2): 23-38 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • MANTHEY, Ulrich & Wolfgang DENZER 1991. Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae) II. Allgemeine Angaben zur Biologie und Terraristik. Sauria 13 (2): 19-22 - get paper here
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Thireau, Michel;Sprackland, Robert George; Sprackland, Teri 1998. A report on Seba's specimens in the herpetological collection of the Muséum National d'Histoire Naturelle, Paris, and their status as Linnaean types. The Linnean 13 (4): 38-45 - get paper here
  • Welton, Luke J.;Cameron D. Siler, L. L. Grismer, Arvin C. Diesmos, Jack W. Sites andRafe M. Brown 2016. Archipelago-wide survey of Philippine forest dragons (Agamidae: Gonocephalus): multilocus phylogeny uncovers unprecedented levels of genetic diversity in a biodiversity hotspot. Biological Journal of the Linnean Society, DOI: 10.1111/bij.12878 - get paper here
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