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Gonocephalus doriae PETERS, 1871

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
SubspeciesGonocephalus doriae abbotti COCHRAN 1922
Gonocephalus doriae doriae (PETERS 1871)
Gonocephalus doriae brevis HARVEY et al. 2023 
Common NamesE: Marquis Doria’s Angle-headed Lizard, Peters' Forest Dragon, Doria’s Anglehead Lizard
G: Dorias Winkelkopfagame
E: Abbott’s Anglehead Lizard [abbotti]
G: Abbotts Winkelkopfagame [abbotti] 
SynonymGonyocephalus Doriae PETERS 1871: 570
Gonyocephalus doriae — BOULENGER 1885: 284
Gonyocephalus doriae — DE ROOIJ 1915: 100
Gonocephalus doriae — COCHRAN 1922: 2
Gonocephalus doriae — TWEEDIE 1954
Gonocephalus doriae — MANTHEY & GROSSMANN 1997: 182
Gonocephalus doriae (C. abbotti) — COX et al. 1998: 95
Gonocephalus doriae doriae — MANTHEY & SCHUSTER 1999: 61
Gonocephalus doriae — MANTHEY 2010
Gonocephalus doriae — HARVEY et al. 2023

Gonocephalus doriae abbotti COCHRAN 1922
Gonocephalus abbotti COCHRAN 1922: 1
Goniocephalus abbotti — TAYLOR 1963: 875
Gonocephalus abbotti — WERMUTH 1967: 56
Gonocephalus abbotti — MOODY 1980
Gonocephalus abbotti — WELCH et al. 1990
Gonocephalus doriae abbotti — DENZER & MANTHEY 2009
Gonocephalus doriae abbotti — MANTHEY 2010: 31
Gonocephalus abbotti — GRISMER et al. 2010
Gonocephalus abbotti — GRISMER & QUAH 2019
Gonocephalus doriae abbotti — HARVEY et al. 2023 
DistributionIndonesia (Borneo: Kalimantan)

Type locality: “Sarawak” [Borneo, Malaysia]

abbotti: W Malaysia (Pahang), S Thailand (Pattani, Trang, Yala); Type locality: Trang, Lower Siam (= Thailand)  
TypesSyntypes: ZMB 7147, MSNG 29152 (the ZMB type could not be found by DENZER et al. 1997)
Holotype: MZB 13488, Adult male, Field Tag 19378, Fig. 10) from forest trail on Seulewah Agam mountain, Aceh Besar, Aceh, Sumatra, Indonesia, 5.45318N, 95.675048E, 993 m elevation, collected at 2302 h, on 1 August 2015 by M. B. Harvey, E. Wostl, and M. Iksan. Paratypes (4).—Unsexed juvenile (MZB 13485) and adult female (UTA 66101) both from 685–971 m along forested trail at Seulewah Agam, collected 30 July 2015 by M. B. Harvey, E. Wostl, I. Fonna, E. N. Smith, G. C. Sarker, F. Akhsani, P. Thammachoti, Syaripudin, M. Ikshan. Unsexed juvenile from Gunung Batee Meucica, Aceh Besar, Aceh, Sumatra, Indonesia, 5.266078N, 95.539928E, 341 m, collected 1 August 2015 by I. Sidik, I. Fonna, P. Thammachoti, G. C. Sarker, E. N. Smith, and Syaripudin. Unsexed juvenile (UTA 66102, GenBank accession number OP070039) from near road leading from Takengon to Beutong, Aceh, Sumatra, Indonesia, 4.470468N, 96.545388E, 1023 m, collected 5 August 2015 by E. N. Smith, G. C. Sarker, and F. Akhsani. [brevis]
Holotype: USNM 24028, adult female [abbotti] 
CommentSynonymy: Gonocephalus doriae abbotti COCHRAN 1922 has been treated as valid species by most recent authors. However, Harvey et al. 2023 suggested to treat it as subspecies of doriae. 
EtymologyNamed after Marchese Giacomo Doria (1840-1913), Italian zoologist who collected in Persia (Iran) with de Filippi (1862-1863) and in Borneo with Beccari (1865-1866). He founded the Museo Civico di Storia Naturale, Turin (1867-1913), and was its first Director.

G. d. brevis is named after the Latin masculine adjective meaning short. Gonocephalus doriae brevis has the shortest tail of any angle-head and this trait usually distinguishes this subspecies from both G. d. abbotti and G. d. doriae.

G. d. abbotti was named after its discoverer, Dr. W. L. Abbott. 
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Cochran, D. M. 1922. Description of a new species of agamid lizard from the Malay Peninsula. Proc. US Natl. Mus. 60: 1-3 - get paper here
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Denzer, W. & Manthey, U. 2009. Remarks on the type specimen of Gonocephalus mjobergi Smith, 1925 (Sauria: Agamidae). Bonner zoologische Beiträge 56 (4): 255–258 - get paper here
  • Evers, M. 2013. Der Traum vom Urwald im Wohnzimmer. Winkelkopfagamen (Gonocephalus) im Terrarium. Reptilia (Münster) 18 (102): 50-59 - get paper here
  • Evers, Michael 2010. Winkelkopfagamen. Gonocephalus belli, C. chamaeleontinus, G. doriae abbotti. Natur und Tier Verlag, 64 pp. - get paper here
  • Goldberg, Stephen R. and L. Lee Grismer. 2014. Gonocephalus abbotti (Abbott's anglehead lizard) reproduction. Herpetological Review 45 (4): 696 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L. Lee; Chan K. Onn, Jesse L. Grismer, Perry L. Wood, Jr., and A. Norhayati 2010. A CHECKLIST OF THE HERPETOFAUNA OF THE BANJARAN BINTANG, PENINSULAR MALAYSIA. Russ. J. Herpetol. 17 (2): 147-160 - get paper here
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  • Grossmann, W. & tillack, F. 2004. Gonocephalus doriae abbotti COCHRAN 1922. Sauria, Berlin 26 (3): 2 - get paper here
  • Harvey, Michael B., Sarker, Goutam C., Sidik, Irvan, Kurniawan, Nia, and Smith, Eric N. 2023. A Contribution to the Systematics of Sunda Shelf Angle- Headed Dragons (Agamidae: Gonocephalus) with the Description of New Taxa from Sumatra. Herpetological Monographs, 37(1): 1-40 - get paper here
  • HÖLßIG, C. 2012. Zur Haltung und Vermehrung von Gonocephalus doriae abbotti COCHRAN, 1922. Sauria 34 (1): 43-50 - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Manthey, U. & DENZER, W. 1993. Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae) VI. Die chamaeleontinus-Gruppe. Sauria 15 (2): 23-38 - get paper here
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  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • Peters, W. 1871. Uber neue Reptilien aus Ostafrika und Sarawak (Borneo), vorzüglich aus der Sammlung des Hrn. Marquis J. Doria zu Genua. Mber. k. preuss. Akad. Wiss., Berlin [1871]: 566-581 - get paper here
  • Sumarli, Alex X.; L. Lee Grismer, Shahrul Anuar, Mohd Abdul Muin and Evan S.H. Quah 2015. First report on the amphibians and reptiles of a remote mountain, Gunung Tebu in northeastern Peninsular Malaysia. Check List 11 (4): 1679 - get paper here
  • Taylor, E.H. 1963. The lizards of Thailand. Univ. Kansas Sci. Bull. 44: 687-1077. - get paper here
  • Tweedie, M.W.F. 1954. Notes on Malayan reptiles, No.3. Bull. Raffles Mus. No 25: 107-117
  • UJANG MUHAMMAD AINI, HADIMAR; KARMELE LLANO SÁNCHEZ, MARIUS JOSCHA MAIWALD & INDRANEIL DAS. 2021. Geographic distribution: GONOCEPHALUS DORIAE (Marquis Doria’s Angle-headed Lizard). INDONESIA: WEST KALIMANTAN PROVINCE: Kayong Utara Regency. Herpetological Review 52 (3): 576–577. - get paper here
  • Welton, Luke J.;Cameron D. Siler, L. L. Grismer, Arvin C. Diesmos, Jack W. Sites andRafe M. Brown 2016. Archipelago-wide survey of Philippine forest dragons (Agamidae: Gonocephalus): multilocus phylogeny uncovers unprecedented levels of genetic diversity in a biodiversity hotspot. Biological Journal of the Linnean Society, DOI: 10.1111/bij.12878 - get paper here
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