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Hebius arquus (DAVID & VOGEL, 2010)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common Names 
SynonymAmphiesma arquus DAVID & VOGEL 2010
Hebius arquus — GUO et al. 2014
Amphiesma arquus — WALLACH et al. 2014: 28 
DistributionIndonesia (Borneo)

Type locality: “Borneo”  
Reproduction 
TypesHolotype: NMW 37943, an adult male. Deposited by Franz Steindachner, 1878. 
CommentAmphiesma arquus sp. nov. is distinguished from other species of the Sunda Region by the combination of an elongated body, 17 + 3 enlarged maxillary teeth, 17 – 17 – 15 dorsal scale rows, all strongly keeled, absence of a loreal, a high number of ventrals and subcaudals, supralabials ornate with white arched markings, and an obscured venter.

Diagnosis. A species of the genus Amphiesma, characterized by a combination of the following characters: (1) elongate body; (2) a long tail, amounting for 30% of the total length; (3) 17 gradually enlarged maxillary teeth, followed without diastema by 3 moderately enlarged posterior teeth; (4) nostrils slightly dorsolateral; (5) 17 – 17 – 15 DSR, all strongly keeled; (6) no loreal, prefrontal in contact with SL 2 – 3; (7) a high number of ventrals (187); (8) supralabials dark, the first seven supralabials ornate on their upper part by an arched white streak; and (9) a largely obscured venter. A comparison with other species of the genus Amphiesma and of other natricine genera of Borneo and adjacent regions is given in the Discussion. The pattern of supralabials, the high number of ventral scales and the lack of loreal scales are enough to distinguish Amphiesma arquus sp. nov. from all other natricine species. 
EtymologyThe specific nomen is the Latin noun arquus (-us), meaning “arches,” by reference to the white arched streaks on the upper part of each anterior supralabial. It is here considered a noun in apposition in the plural, and not an adjective. 
References
  • David, Patrick and Gernot Vogel. 2010. A new species of Natricine snake genus Amphiesma from Borneo (Squamata: Natricidae). Russ. J. Herpetol. 17 (2): 121-127 - get paper here
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
 
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