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Hebius atemporale (BOURRET, 1934)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Common NamesE: Tonkin Keelback
Chinese: 无颞鳞腹链蛇 
SynonymNatrix atemporalis BOURRET 1934
Natrix atemporalis — SMITH 1943: 287
Amphiesma atemporalis — MALNATE 1960
Amphiesma atemporalis — ZHAO 2006
Amphiesma atemporale — DAVID et al. 2007
Hebius atemporale — GUO et al. 2014
Amphiesma atemporale — WALLACH et al. 2014: 28
Hebius atemporalis — ZHOU et al. 2019 
DistributionN Vietnam,
China (Hong Kong, Guangdong, and Guangxi west to Yunnan and Guizhou)

Type locality: Ton-king (= North Vietnam)  
TypesSyntypes: MNHN-RA 1935.0077-0078, MNHN-RA 1935.0450, MNHN-RA 1938.0119-0120 (5 syntypes); perhaps MNHN-RA 1958.0461-0462 are also syntypes? 
  • Bourret, RENÉ. 1934. Notes herpétologiques sur l'Indochine Française VI-VII. Bull. Gén. Instr. Publique, Hanoi, 1934 (4): 73-84
  • David, P., Bain, R. H., Nguyen Quang Truong, Orlov, N. L., Vogel, G., Vu Ngoc Thanh & T. Ziegler 2007. A new species of the natricine snake genus Amphiesma from the Indochinese Region (Squamata: Colubridae: Natricinae). Zootaxa 1462: 41-60 - get paper here
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Malnate, E.V. & Romer, J.D. 1969. A snake new to the fauna of Hong Kong and China: Amphiesma atemporalis (Bourret). Notulae Naturae, Acad. Nat. Sci. Philadelphia, 424, 1–8.
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp.
  • Ziegler, T., and Quyet, L.K. 2006. A new natricine snake of the genus Amphiesma (Squamata: Colubridae: Natricinae) from the central Truong Son, Vietnam. Zootaxa 1225:39-56 - get paper here
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