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Hebius deschauenseei (TAYLOR, 1934)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Northern Keelback, Deschauensee’s Keelback
G: Deschauensees Wassernatter 
SynonymNatrix deschauenseei TAYLOR 1934: 300
Macropophis deschauenseei — BOURRET 1936: 105
Amphiesma deschauenseei — MALNATE 1960: 50
Amphiesma modesta — NABHITABHATA1987: 42
Amphiesma deschauenseei — DAVID, VOGEL & PAUWELS 1998
Amphiesma deschauenseei — DAVID et al. 2007
Hebius deschauenseei — GUO et al. 2014
Amphiesma deschauenseei — WALLACH et al. 2014: 29
Amphiesma deschauenseei — CHAN-ARD et al. 2015: 230
Hebius deschauenseei — DAVID et al. 2021 
DistributionThailand (Chiang Mai; Chiang Rai; Uthai Thani)

Type locality: Type loc. Chieng Mai, North Siam.  
TypesHolotype: ANSP 19927, Acad. Nat. Sci. Philadelphia. Coll., Baron R. M. de Schauensee, between Jan. 1 — Mar. 1, 1933. 
DiagnosisDiagnosis: A moderately to large sized species of the genus Hebius characterized by the combination of (1) 19-19-17 dorsal scale rows, moderately or weakly keeled at midbody, more strongly keeled posteriorly excepted scales of the 1st DSR, all smooth; (2) dorsal scales around the base of the tail weakly keeled; (3) head moderately distinct from the neck; (4) eye rather large; (5) 29–30 maxillary teeth, the last two moderately enlarged; (6) tail long and tapering, with a ratio TaL/TL at least equal to 0.33; (7) VEN 149–168; (8) SC 115–141; (9) prefrontal 2; (10) usually 5th–6th SL entering orbit; (11) 1 anterior temporal; (12) venter never entirely dark: along its whole length, creamish-yellow or pale yellowish-grey with three parallel rows of dark blotches, one on the outer part of each ventral, and one on the central part of the venter producing three irregular, parallel stripes separated from each other by a narrow, pale area; (13) dorsum and sides olive-brown, olive-grey or greyish-brown, sometimes dark brown; (14) dorsal surface with irregular, blackish-brown or very dark grey blotches, not producing a chequered pattern; (15) a dorsolateral series of large, elongate blotches, yellow-ochre, rusty red or yellowish-brown, enlarged on the first quarter to third of the body, progressively smaller and united, forming a dorsolateral stripe, usually extending along the whole of the body on the 4th–5th or 5th–6th dorsal scale rows; (16) postocular streak present, not visible in dark specimens; and (17) an ochre-yellow or yellowish-brown streak on each side of the neck and nape forming an incomplete collar (David et al. 2021). 
CommentListed as synonym of Natrix (= Amphiesma) modesta by SMITH 1943: 290.

Distribution: not in Vietnam, fide P. David, pers. comm., 21 Apr 2021. 
EtymologyNamed after Dr. Rodolphe Mayer Deschauensee (1901-1984), former Curator and Director of the Academy of Natural Sciences, Philadelphia and among the collectors of the holotype. 
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • David P., G. Vogel & Pauwels,O. 1998. Amphiesma optatum (Hu & Djao, 1966) (Serpentes, Colubridae): an addition to the snake fauna of Vietnam, with a list of the species of the genus Amphiesma and a note on its type species. Journal of Taiwan Museum 51 (2): 83-92 - get paper here
  • David, P., Bain, R. H., Nguyen Quang Truong, Orlov, N. L., Vogel, G., Vu Ngoc Thanh & T. Ziegler 2007. A new species of the natricine snake genus Amphiesma from the Indochinese Region (Squamata: Colubridae: Natricinae). Zootaxa 1462: 41-60 - get paper here
  • DAVID, PATRICK; GERNOT VOGEL, TRUONG QUANG NGUYEN, NIKOLAI L. ORLOV, OLIVIER S. G. PAUWELS, ALEXANDRE TEYNIÉ, THOMAS ZIEGLER 2021. A revision of the dark-bellied, stream-dwelling snakes of the genus Hebius (Reptilia: Squamata: Natricidae) with the description of a new species from China, Vietnam and Thailand. Zootaxa 4911: 1–61 - get paper here
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Taylor, E.H. 1944. Present location of certain herpetological and other type specimens. Univ. Kansas Sci. Bull. 30 (1): 117-187. - get paper here
  • Taylor,E.H. 1934. Zoological results of the third De Schauensee Siamese Expedition, part IlI. Amphibians and Reptiles. Proc. Acad. Nat. Sci. Philad. 86: 281-310 - get paper here
  • Taylor,E.H. 1965. The serpents of Thailand and adjacent waters. Univ. Kansas Sci. Bull. 45 (9): 609-1096 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Ziegler, T., and Quyet, L.K. 2006. A new natricine snake of the genus Amphiesma (Squamata: Colubridae: Natricinae) from the central Truong Son, Vietnam. Zootaxa 1225: 39-56 - get paper here
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