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Hebius ishigakiensis (MALNATE & MUNSTERMAN, 1960)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Yaeyama keelback 
SynonymNatrix pryeri ishigakiense MALNATE & MUNSTERMAN 1960: 61
Tropidonotus pryeri — WALL 1905 : 515 (part.)
Natrix pryeri — STEJNEGER 1907: 284 (part.)
Amphiesma pryeri — MALNATE 1960: 51 (part.)
Amphiesma pryeri ishigakiense — MALNATE 1962
Amphiesma pryeri ishigakiense — MATSUI 1996
Amphiesma ishigakiense — OTA and IWANAGA 1997
Amphiesma pryeri ishigakiense — GORIS & MAEDA 2004: 208
Hebius ishigakiense — GUO et al. 2014
Amphiesma ishigakiense — WALLACH et al. 2014: 30
Hebius ishigakiensis — TAKEUCHI 2018 
DistributionJapan (Ryukyu Islands)

Type locality: Ishigaki-shima, Yaeyama group, Riukiu Islands.  
TypesHolotype: CAS 21913 
DiagnosisDiagnosis: Ota & Iwanaga 1997 
CommentRaised to species status by Ota & Iwanaga 1997. 
Etymologynamed after the type locality. 
  • Blackburn, Daniel G. 1999. Are Viviparity and Egg-guarding Evolutionarily Labile in Squamates? Herpetologica 55 (4): 556-573 - get paper here
  • Goris, R.C. & Maeda, N. 2004. Guide to the Amphibians and Reptiles of Japan. Krieger, Malabar, 285 pp.
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Kaito, Takuma; and Mamoru Toda 2016. The biogeographical history of Asian keelback snakes of the genus Hebius (Squamata: Colubridae: Natricinae) in the Ryukyu Archipelago, Japan. Biological Journal of the Linnean Society, DOI: 10.1111/bij.12726 - get paper here
  • Malnate, E.V. 1962. The relationships of five species of the Asiatic natricine snake genus Amphiesma. Proc. Acad. Nat. Sci. Philadelphia, 114, 251–299. - get paper here
  • Malnate, E.V. and MUNSTERMAN,H.E. 1960. Interpopulation variation in the colubrid snake Natrix pryeri from the Riukiu Islands, with description of a new subspecies. Proc. Cal. Acad. Sci. (4) 31 (2): 51-67 - get paper here
  • Ota, HIDETOSHI & SETSUKO IWANAGA 1997. A systematic review of the snakes allied to Amphiesma pryeri (Boulenger) (Squamata: Colubridae) in the Ryukyu Archipelago, Japan. Zoological Journal of the Linnean Society 121 (3): 339-360 - get paper here
  • Takeuchi, H; 2018. Hebius ishigakiensis (Yaeyama Keelback) Diet. Herpetological Review 49 (4): 753 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Ziegler, T., and Quyet, L.K. 2006. A new natricine snake of the genus Amphiesma (Squamata: Colubridae: Natricinae) from the central Truong Son, Vietnam. Zootaxa 1225: 39-56 - get paper here
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