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Hebius leucomystax (DAVID, BAIN, QUANG TRUONG, ORLOV, VOGEL, NGOC THANH & ZIEGLER, 2007)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesE: White-lipped Keelback
F: Amphiesme à lêvres blanches
G: Weißlippen-Gebirgswassernatter
V: Ran sai mep trang (Vietnamese) 
SynonymAmphiesma leucomystax DAVID, BAIN, QUANG TRUONG, ORLOV, VOGEL, NGOC THANH & ZIEGLER 2007
Hebius leucomystax — GUO et al. 2014
Amphiesma leucomystax — PAUWELS et al. 2015
Amphiesma leucomystax — WALLACH et al. 2014: 31 
DistributionC Vietnam [(Quy Chau District), Ha Tinh (Cam Xuyen and
Huong Son Districts), Quang Binh (Bo Trach and Minh Hoa Districts), Quang Tri (A Luoi, Phu Loc and Huong Hoa Districts), Thua Thien-Hue (Huong Thuy and A Luoi Districts), Quang Nam (Ba Na National Park, Hoa Vang District), Kon Tum (Ngoc Linh Mt., on the border between Quang Nam and Kon Tum Provinces), and Gia Lai (An Khe District)], Thailand, Laos

Type locality: southeastern border of the Ky Anh - Ke Go lowland forest protected area, surroundings 18°00’N-106°06’E, Cam Xuyen District, Ha Tinh Province, Vietnam, 125 m elevation. Map legend:
Type locality - Type locality.
 
Reproductionoviparous 
TypesHolotype: ZFMK 71702, an adult female. Coll. by Thomas Ziegler, 12 July1997. 
CommentDiagnosis: A species of the genus Amphiesma, characterized by a combination of the following characters: (1) a broad, continuous, white stripe extending under the eye from the tip of the snout across the upper half of supralabials and the nape to produce a V-like chevron; (2) vertically elongated or divided dorsolateral spots, salmon or rusty red (cream in preservative) on a dark grey background; (3) tips of ventrals black, with additional faint and ill defined dark blotches on inner side; (4) 19-19-17 DSR, distinctly keeled on rows 3–1 or 4–1, smooth on rows 1–2 or 1–3; scales of 1st DSR enlarged; (5) 154–166 ventral plates; (6) internasals narrowed anteriorly; (7) 1 anterior temporal, 1 preocular and usually 3 postoculars; (8) a moderately sized eye. 
EtymologyEtymology.—The specific nomen derives from the Greek adjective , leucos, meaning white, and the Greek noun “mystax”, meaning either a moustache or an upper lip, a reference to the broad white stripe extending on the upper lips of this species. 
References
  • David, P., Bain, R. H., Nguyen Quang Truong, Orlov, N. L., Vogel, G., Vu Ngoc Thanh & T. Ziegler 2007. A new species of the natricine snake genus Amphiesma from the Indochinese Region (Squamata: Colubridae: Natricinae). Zootaxa 1462: 41-60 - get paper here
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Nguyen, Luan Thanh; Ha Van Hoang, Thang Tai Nguyen,<br />McCormack Timothy E.M., Sang Ngoc Nguyen 2016. A COLLECTION OF AMPHIBIANS AND REPTILES FROM BAC HUONG HOA NATURE RESERVE, QUANG TRI PROVINCE, VIETNAM - get paper here
  • Ostroshabov, Alexander A., Nikolai L. Orlov and Larissa K. Iohanssen. 2016. Hebius leucomystax (White-lipped Keelback) Diet and foraging behavior. Herpetological Review 47 (2): 311-312
  • Pauwels, Olivier S. G.; Patrick David, Tanya Chan-ard 2015. First Confirmed Record of the Stream-Dwelling Snake Amphiesma leucomystax (Squamata: Natricidae) in Thailand. Russ. J. Herpetol. 22 (2): 136-138 - get paper here
  • Sang, Nguyen Van; Ho Thu Cuc, Nguyen, Quang Truong 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Stuart, B.L. & Heatwole, H. 2008. Country records of snakes from Laos. Hamadryad 33: 97–106 - get paper here
  • Stuart, B.L., Rowley, J.J.L., Neang, T., Emmett, D.A. & Sitha, Som 2010. Significant new records of amphibians and reptiles from Virachey National Park, northeastern Cambodia. Cambodian Journal of Natural History 2010(1): 38–47 - get paper here
  • Thompson, C. & Thompson, T. 2008. First contact in the Greater Mekong - new species discoveries. WWF, 40 pp. - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
 
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