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Hebius pryeri (BOULENGER, 1887)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesPryer's Keelback 
SynonymTropidonotus pryeri BOULENGER 1887: 149
Tropidonotus pryeri — BOULENGER 1893: 250
Natrix pryeri - STEJNEGER 1907: 284
Natrix pryeri — SCLATER 1950
Amphiesma pryeri - MALNATE & MUNSTERMANN 1960: 51
Hebius pryeri — GUO et al. 2014
Amphiesma pryeri — WALLACH et al. 2014: 32 
DistributionJapan (Ryukyu Islands)

Type locality: Ishigaki-shima, Yaeyama group, Riukiu Islands  
Reproductionoviparous 
TypesHolotype: CAS 21913
Paratypes: CAS 21914, 21917, 21919, males; CAS 21912, 21915, 21916, 21918, 21921 (all collected on Ishigaki-shima, Yaeyama group, Riukiu Islands) 
CommentSome subspecies of A. pryeri have been elevated to full species status, e.g. A.p.ishigakiense.

Diagnosis: Ota & Iwanaga 1997 
References
  • Ackermann, L. 2016. In den Wäldern von Okinawa. Terraria-Elaphe 2016 (5): 48-51 - get paper here
  • Boulenger, G.A. 1887. On a collection of reptiles and batrachians made by Mr. H. Pryer in the Loo Choo Islands. Proc. Zool. Soc. London 1887: 146-150 - get paper here
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Goris, R.C. & Maeda, N. 2004. Guide to the Amphibians and Reptiles of Japan. Krieger, Malabar, 285 pp.
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Kaito, Takuma; and Mamoru Toda 2016. The biogeographical history of Asian keelback snakes of the genus Hebius (Squamata: Colubridae: Natricinae) in the Ryukyu Archipelago, Japan. Biological Journal of the Linnean Society, DOI: 10.1111/bij.12726 - get paper here
  • Mori,M. 1982. Japans Schlangen, Vols. 1-3 (80, 102, 123 pp.). Tokyo (Igaku-Shoin Ltd.) [bilingual edition in German and Japanese]
  • Ota, HIDETOSHI & SETSUKO IWANAGA 1997. A systematic review of the snakes allied to Amphiesma pryeri (Boulenger) (Squamata: Colubridae) in the Ryukyu Archipelago, Japan. Zoological Journal of the Linnean Society 121 (3): 339-360
  • Sclater, J.A. 1950. Bionomic notes on some reptiles form Okinawa Shima, Ryukyu Islands, Japan. Ann. Mag. Nat. Hist. (12) 3: 113-116
  • Stejneger, LEONHARD H. 1907. Herpetology of Japan and adjacent territory. Bull. US Natl. Mus. 58: xx, 1-577 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
  • Ziegler, T., and Quyet, L.K. 2006. A new natricine snake of the genus Amphiesma (Squamata: Colubridae: Natricinae) from the central Truong Son, Vietnam. Zootaxa 1225:39-56 - get paper here
 
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