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Helicops angulatus (LINNAEUS, 1758)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesBrown-banded watersnake, Mountain Keelback 
SynonymColuber angulatus LINNAEUS 1758: 217
Coluber alidras LINNAEUS 1758
Coluber Surinamensis SHAW 1802: 460
Natrix aspera WAGLER 1824: 37
Helicops angulatus — WAGLER 1830: 171
Helicops angulatus — DUMÉRIL, BIBRON & DUMÉRIL 1854: 746
Helicops angulatus — GÜNTHER 1861: 428
Helicops fumigatus COPE 1868: 308
Helicops cyclops COPE 1868: 309
Uranops angulatus — SCLATER 1891: 45
Helicops angulatus — BOULENGER 1893: 278
Helicops angulata — BEEBE 1946: 28
Helicops angulatus — PETERS & OREJAS-MIRANDA 1970: 123
Helicops angulatus — GASC & RODRIGUES 1980
Helicops angulatus — GORZULA & SEÑARIS 1999
Helicops angulatus — KAWASHITA-RIBEIRO et al. 2013
Helicops angulatus — WALLACH et al. 2014: 321 
DistributionVenezuela (Amazonas, Apure, Bolivar, Monagas, Delta Amacuro, Sucre, Portuguesa, Anzoátegui, Guárico, Cojedes), Colombia, Brazil (Pará, Rondonia, Goias, Mato Grosso, Sergipe, S Ceará, Acre, Bahia etc.), Bolivia, Peru, Trinidad, Ecuador, French Guiana, Guyana

Type locality: “Asia” (in error) Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
ReproductionHelicops angulatus is only one of 3 or so species that are known to have both viviparous and oviparous reproduction (Rossman 1974, 1984, Braz et al. 2016) 
TypesHolotype: NRM (also given as RMS)
Holotype: NRM [Coluber alidras]
Holotype: ANSP 5183; Brazil, Bahia [Helicops cyclops]
Holotype: ANSP 5132; Surinam; [Helicops fumigatus]
Lectotype: ZSM 1528/0, female [Natrix aspera] 
CommentSynonymy partly after PETERS & OREJAS-MIRANDA 1970. 
References
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