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Hemidactylus gubanensis MAZUCH, JANÁK, VELENSKÁ, NISTRI, ABDIRAHMAN-ELMI & ŠMÍD, 2024

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Guban gecko
Somali: Qoroto [pronounced Koroto] 
SynonymHemidactylus gubanensis MAZUCH, JANÁK, VELENSKÁ, NISTRI, ABDIRAHMAN ELMI & ŠMÍD 2024: 8
Hemidactylus sinaitus — PARKER 1942
Hemidactylus sinaitus — LANZA 1983, 1990
Hemidactylus sinaitus — LARGEN & SPAWLS 2006 (in part)
Hemidactylus sinaitus — LARGEN & SPAWLS 2010 (in part)
Hemidactylus turcicus turcicus — LOVERIDGE 1947 (in part)
Hemidactylus afarensis — ŠMÍD et al. 2020 (in part)
Hemidactylus afarensis — SPAWLS et al. 2023 (in part) 
DistributionSomaliland

Type locality: Somaliland, Sahil Region, 1 km west of Girigoyan (or Gari Goan, Gorigaan) Mountain (10.297° N, 44.996° E; datum WGS84 hereafter, 261 m above sea level [asl])  
Reproduction 
TypesHolotype: NMP-P6V 76683 (sample code TMHC_998), adult male, collected by T. Mazuch on 27 January 2022 (Figures 4, 5, 6a).
Paratypes (four): NMP-P6V 76682 (sample code TMHC_997), adult female, NMP-P6V 76684 (sample code TMHC_999), adult male; same collection data as the holotype. NMP-P6V 76680 (sample code S22-03), juvenile (Figure 6b), NMP-P6V 76681 (sample code TMHC_996), adult male; Somaliland, Sahil Region, 1 km north-west of Girigoyan (or Gari Goan) Mountain (10.309° N, 45.0° E, 241 m asl), collected by T. Mazuch on 25 January 2022.
Other material (16): TMHC 2024.01.1036, TMHC 2024.01.1038-1039, adult males, TMHC 2024.01.1037, adult female; Somaliland, Awdal Region, 3.5 km north-west of Jidhi village (10.636° N, 43.043° E, 468 m asl), collected by T. Mazuch, D. Hlaváč, P. Konečný, H. Sh. A. Elmi on 15 November 2023. TMHC 2024.01.1040–1041, adult females; Somaliland, Awdal Region, 1 km north-west of Gargara Galbeed (10.774° N, 43.031° E, 350 m asl), collected by T. Mazuch, D. Hlaváč, P. Konečný, H. Sh. A. Elmi on 15 November 2023. TMHC 2024.01.1042, TMHC 2024.01.1044, TMHC 2024.01.1047, adult males, TMHC 2024.01.1043, TMHC 2024.01.1045–1046, adult females; Somaliland, Sahil Region, 6 km east-southeast of Xagal village, (10.235° N, 45.793° E, 185 m asl), collected by T. Mazuch, D. Hlaváč, P. Konečný, H. Sh. A. Elmi on 22 November 2023. TMHC 2017.12.872-873, adult females (Figure 6c); Somaliland, Awdal Region, Geerissa village (10.629° N, 43.441° E, 223 m asl), collected by T. Mazuch on 11 September 2017. NHMUK 1937.12.5.293, adult male; Somaliland, Awdal Region, ca. 25 km south-western of Geerissa (10.5° N, 43.25° E, 485 m asl), collected by R.H.R. Taylor on 18 April 1933. NHMUK 1937.12.5.294, juvenile male; Somaliland, Awdal Region, ca. 25 km west of Zeila (11.416667° N, 43.25° E, 29 m asl), collected by R.H.R. Taylor on 9 January 1934. The NHMUK specimens were compared based on photographs but were not included in the morphological analysis. 
DiagnosisDiagnosis: A species of the African radiation of the arid clade of Hemidactylus (sensu Šmíd et al. 2013a, 2020) with the following combination of morphological traits: (1) small size, SVL 41.5–53.3 mm in males, 36.5–52.9 mm in females; (2) large and robust head, head depth 32–50% of head length, head width 58–82% of head length; (3) relatively short tail, tail length 94.0–111.3 % of SVL; (4) uppermost nasals in broad contact; (5) large anterior postmentals usually in wide contact (in 84% specimens) or in one point (16% specimens); (6) large anterior postmentals usually in contact with the first and second infralabials (in 82% specimens, with the first infralabial only in 18% specimens); (7) 7–10 supralabials; (8) 6–8 infralabials; (9) dorsum with 14–16 rows of enlarged, strongly keeled, conical tubercles; (10) other dorsal scales smooth, juxtaposed, and rather homogeneous; (11) 3–6 (counted longitudinally) and 2–3 (transversely) small smooth dorsal scales between dorsal tubercles; (12) 22–27 tail segments; (13) 2–4 rows of small scales between tail tubercles of two adjacent tail segments; (14) subcaudals enlarged; (15) rather long free distal phalanges of digits (relative length of free distal phalanges and proximal parts with lamellae on the 3rd toe 0.35–0.57, mean 0.45 ± 0.05); (16) 5–6 lamellae under the 1st finger, 7–8 lamellae under the 4th finger, 5–6 lamellae under the 1st toe, and 8–11 lamellae under the 4th toe; (17) 4–7 precloacal pores in males; (18) presence of lateral postanal tubercles; (19) in life, body colour light beige-brownish with irregular and indistinct dark mottling, some tubercles whitish and some dark; tail with similar colouration and pattern like body or with very slightly marked light and dark irregular bands especially in the anterior half; head paler with darker brown snout and rather irregular pale brownish-yellow stripe running from posterior margin of eye to above the ear (Figure 6a,b,c). (Mazuch et al. 2024)


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Comment 
EtymologyNamed after Guban, a hot coastal plain in northern Somaliland where all specimens have been collected and where it is one of the most abundant ground-dwelling gecko species. The word Guban translates as ‘burnt’ in Somali language, referring to the torrid weather conditions that prevail in the region throughout most of the year (March−September). 
References
  • Mazuch, T., Janák, V., Velenská, D., Nistri, A., Elmi, H. S. A., & Šmíd, J. 2024. Two new species of Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from the coastal areas of northern Somaliland. African Zoology, 1-24 - get paper here
 
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