Hemidactylus ulii ŠMÍD, MORAVEC, KRATOCHVÍL, GVOŽDÍK, NASHER, BUSAIS, WILMS, SHOBRAK, CARRANZA, 2013
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Hemidactylus ulii
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| Higher Taxa | Gekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) |
| Subspecies | |
| Common Names | |
| Synonym | Hemidactylus ulii ŠMÍD, MORAVEC, KRATOCHVÍL, GVOžDÍK, NASHER, BUSAIS, WILMS, SHOBRAK, CARRANZA 2013 Hemidactylus turcicus – RÖSLER & WRNAIK 1998: 120 (part.). Hemidactylus sp. ‘OTU7’ – BUSAIS & JOGER 2011a: 27 Hemidactylus sp. ‘OTU7’ – BUSAIS & JOGER 2011b: 268 Hemidactylus sp. ‘OTU7’ – CARRANZA & ARNOLD 2012: 95 Hemidactylus sp. 4 – MORAVEC et al. 2011: 25 Hemidactylus sp. 4 – ŠMÍD et al. 2013: 3 |
| Distribution | SW Yemen Type locality: Yemen, Ta’izz governorate, Al Hababi (13.333°N, 43.722°E), 463 m elevation. |
| Reproduction | oviparous (manual and phylogenetic imputation, fide Zimin et al. 2022) |
| Types | Holotype: NMP P6V 74833/2, adult male (MorphoBank M305892–M305902), collected by L. Kratochvíl, 28. X. 2007. Paratypes. NMP6V 74833/1 (adult male, MorphoBank M305884–M305891), same collecting data as holotype; NMP6V 74831/1–2 (one adult and one subadult female, MorphoBank M305854–M305863, M305864–M305870), Yemen, Abyan governorate, Al Hadr (13.877°N, 45.8°E), 1151 m a.s.l., collected by L. Kratochvíl on 22. X. 2005; NMP6V 74832/1–2 (two subadult females, MorphoBank M305871– M305875, M305876–M305883), Yemen, Ta’izz governorate, ca. 3 km S of Najd an Nashamah by road (13.358°N, 43.957°E), 1182 m a.s.l., collected by L. Kratochvíl on 26. X. 2007; NMP6V 74834/1–2 (one adult and one subadult female, Morpho- Bank M305903–M305911), Yemen, Dhamar governorate, Wadi Zabid (14.147°N, 43.517°E), 292 m a.s.l., collected by L. Kratochvíl on 29. X. 2007; NHM-BS N41916 (juvenile, MorphoBank M305842–M305852), Yemen, Al Bayda’ governorate, Rad- man (14.1°N, 45.283°E), collected by W. Mustafa on 13. XI. 2007. |
| Diagnosis | Diagnosis: A small species of the ‘Hemidactylus saba species group’ within the Arabian radiation of the Arid clade of Hemidactylus, as evidenced by the mtDNA and nDNA analyses. The new species is characterized by the following combination of molecular and morphological characters: (1) Uncorrected genetic distances from H. saba: 9.9–10.7% in 12S, 13.5–14.9% in cytb; from H. granosus: 10.2–12.3% in 12S, 11.2– 13.5% in cytb; (2) small size with a maximum recorded SVL 40.7 mm (36.8–40.4 mm in males, 39.4–40.7 mm in females); (3) moderately robust head, head length 28–30% of SVL, head width 70–75% of head length, head depth 37–46% of head length; (4) tail length 116% of SVL (only 1 specimen with intact tail); (5) uppermost nasals separated by a small shield (60% specimens) or in wide contact (40%); (6) large anterior postmentals in wide mutual contact in 90% of individuals, and in contact with the 1st and 2nd lower labial (scarcely and unilaterally with the 1st lower labial only); (7) 8–10 upper labials; (8) 7–9 lower labials; (9) dorsum with 12-16 longitudinal rows of enlarged, slightly keeled, conical tubercles; (10) 5–6 lamellae under the 1st toe and 8–9 lamellae under the 4th toe; (11) ca. 6–8 tail segments bearing 6 tubercles; (12) 8 preanal pores in one continuous row in males; (13) subcaudals enlarged; (14) in alcohol dorsum brownish grey with a pattern of more or less conspicuous dark transverse bands starting on the nape, tail with 9 dark brown transverse bands. |
| Comment | Distribution: see map in ŠMÍD et al. 2016: Fig. 3. Sympatry: Bunopus spatalurus Anderson, 1901; Hemidactylus y. yerburii Anderson, 1895; Pristurus crucifer (Valenciennes, 1861); P. flavipunctatus Rüppell, 1835; P. rupestris Blanford, 1874; Ptyodactylus sp.; Tropiocolotes scorteccii Cherchi and Spano, 1963; Acanthodac- tylus sp.; Chamaeleo arabicus Matschie, 1893; Pseudotrapelus sinaitus (Heyden, 1827); Trapelus flavimaculatus Rüppell, 1835; and Pelomedusa subrufa (Bonnaterre, 1789). |
| Etymology | The species epithet “ulii” is a patronym for Prof. Ulrich Joger, a German herpetologist known as Uli among friends, in recognition of his important contribution to the knowledge of the herpetofauna of the Western Palearctic. |
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