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Hemidactylus ulii ŠMÍD, MORAVEC, KRATOCHVÍL, GVOžDÍK, NASHER, BUSAIS, WILMS, SHOBRAK, CARRANZA, 2013

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common Names 
SynonymHemidactylus ulii ŠMÍD, MORAVEC, KRATOCHVÍL, GVOžDÍK, NASHER, BUSAIS, WILMS, SHOBRAK, CARRANZA 2013
Hemidactylus turcicus – RÖSLER & WRNAIK 1998: 120 (part.).
Hemidactylus sp. ‘OTU7’ – BUSAIS & JOGER 2011a: 27
Hemidactylus sp. ‘OTU7’ – BUSAIS & JOGER 2011b: 268
Hemidactylus sp. ‘OTU7’ – CARRANZA & ARNOLD 2012: 95
Hemidactylus sp. 4 – MORAVEC et al. 2011: 25
Hemidactylus sp. 4 – ŠMÍD et al. 2013: 3 
DistributionSW Yemen

Type locality: Yemen, Ta’izz governorate, Al Hababi (13.333°N, 43.722°E), 463 m elevation. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproduction 
TypesHolotype: NMP6V 74833/2, adult male (MorphoBank M305892–M305902), collected by L. Kratochvíl, 28. X. 2007.
Paratypes. NMP6V 74833/1 (adult male, MorphoBank M305884–M305891), same collecting data as holotype; NMP6V 74831/1–2 (one adult and one subadult female, MorphoBank M305854–M305863, M305864–M305870), Yemen, Abyan governorate, Al Hadr (13.877°N, 45.8°E), 1151 m a.s.l., collected by L. Kratochvíl on 22. X. 2005; NMP6V 74832/1–2 (two subadult females, MorphoBank M305871– M305875, M305876–M305883), Yemen, Ta’izz governorate, ca. 3 km S of Najd an Nashamah by road (13.358°N, 43.957°E), 1182 m a.s.l., collected by L. Kratochvíl on 26. X. 2007; NMP6V 74834/1–2 (one adult and one subadult female, Morpho- Bank M305903–M305911), Yemen, Dhamar governorate, Wadi Zabid (14.147°N, 43.517°E), 292 m a.s.l., collected by L. Kratochvíl on 29. X. 2007; NHM-BS N41916 (juvenile, MorphoBank M305842–M305852), Yemen, Al Bayda’ governorate, Rad- man (14.1°N, 45.283°E), collected by W. Mustafa on 13. XI. 2007. 
CommentDiagnosis. A small species of the ‘Hemidactylus saba species group’ within the Arabian radiation of the Arid clade of Hemidactylus, as evidenced by the mtDNA and nDNA analyses. The new species is characterized by the following combination of molecular and morphological characters: (1) Uncorrected genetic distances from H. saba: 9.9–10.7% in 12S, 13.5–14.9% in cytb; from H. granosus: 10.2–12.3% in 12S, 11.2– 13.5% in cytb; (2) small size with a maximum recorded SVL 40.7 mm (36.8–40.4 mm in males, 39.4–40.7 mm in females); (3) moderately robust head, head length 28–30% of SVL, head width 70–75% of head length, head depth 37–46% of head length; (4) tail length 116% of SVL (only 1 specimen with intact tail); (5) uppermost nasals separated by a small shield (60% specimens) or in wide contact (40%); (6) large anterior postmentals in wide mutual contact in 90% of individuals, and in contact with the 1st and 2nd lower labial (scarcely and unilaterally with the 1st lower labial only); (7) 8–10 upper labials; (8) 7–9 lower labials; (9) dorsum with 12-16 longitudinal rows of enlarged, slightly keeled, conical tubercles; (10) 5–6 lamellae under the 1st toe and 8–9 lamellae under the 4th toe; (11) ca. 6–8 tail segments bearing 6 tubercles; (12) 8 preanal pores in one continuous row in males; (13) subcaudals enlarged; (14) in alcohol dorsum brownish grey with a pattern of more or less conspicuous dark transverse bands starting on the nape, tail with 9 dark brown transverse bands.

Distribution: see map in ŠMÍD et al. 2016: Fig. 3.

Sympatry: Bunopus spatalurus Anderson, 1901; Hemidactylus y. yerburii Anderson, 1895; Pristurus crucifer (Valenciennes, 1861); P. flavipunctatus Rüppell, 1835; P. rupestris Blanford, 1874; Ptyodactylus sp.; Tropiocolotes scorteccii Cherchi and Spano, 1963; Acanthodac- tylus sp.; Chamaeleo arabicus Matschie, 1893; Pseudotrapelus sinaitus (Heyden, 1827); Trapelus flavimaculatus Rüppell, 1835; and Pelomedusa subrufa (Bonnaterre, 1789). 
EtymologyThe species epithet “ulii” is a patronym for Prof. Ulrich Joger, a German herpetologist known as Uli among friends, in recognition of his important contribution to the knowledge of the herpetofauna of the Western Palearctic. 
References
  • Rösler, Herbert 2015. Bemerkungen über einige Geckos der Zoologischen Staatssammlung München. Gekkota, Suppl. (2): 3-54
  • Šmíd J, Moravec J, Kratochvíl L, Gvoždík V, Nasher AK, Busais SM, Wilms T, Shobrak MY, Carranza S 2013. Two newly recognized species of Hemidactylus (Squamata, Gekkonidae) from the Arabian Peninsula and Sinai, Egypt. ZooKeys 355: 79–107 - get paper here
  • Šmíd, Jiří; Mohammed Shobrak; Thomas Wilms; Ulrich Joger & Salvador Carranza 2016. Endemic diversification in the mountains: genetic, morphological, and geographical differentiation of the Hemidactylus geckos in southwestern Arabia. Org Divers Evol, DOI 10.1007/s13127-016-0293-3
 
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