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Hemiphyllodactylus cicak COBOS, GRISMER, WOOD, QUAH, ANUAR & MUIN, 2016

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Penang Island Slender Gekko
Malay: Cicak Kerdil Pulau Pinang 
SynonymHemiphyllodactylus cicak COBOS, GRISMER, WOOD, QUAH, ANUAR & MUIN 2016 
DistributionPeninsular Malaysia (Pulau Pinang)

Type locality: 600 m elevation from the living room of the old Ban Hin Lee Guest House on Penang Hill, Pulau Pinang, Peninsular Malaysia (5°25’23.14”N, 100°16’19.79”E)  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: LSUHC (erroneously given as LSUCH in Cobos et al.) 11762, adult male, collected by Evan Quah and L. Lee Grismer on 5 May 2014 at approximately 1800 hrs during a heavy thunderstorm. Paratypes. Three adult females (LSUHC 11763–65) are associated with the same collection data as the holotype. 
DiagnosisDiagnosis: Hemiphyllodactylus cicak sp. nov. can be separated from all other species of Hemiphyllodactylus by possessing the unique combination of characters, having a maximum SVL of 31.4 mm; three circumnasal scales; one or two scales between supranasals (=postrostrals); eight supralabials; eight infralabials; 10 or 11 longitudinally arranged ventral scales at midbody contained within one eye diameter; a series of 42 pore-bearing precloacal-femoral scales in the male; lamellar formula on hand 2–3–3–2; lamellar formula on foot 2–3–3–3; dorsal body pattern consisting of dark, transverse, paravertebral blotches coupled with white speckles; dark pre- and postorbital stripes; faint postorbital stripe extends along the flanks to the hind limbs. These characters, and other diagnostic characters, are scored across all species of Hemiphyllodactylus within the harterti group listed in Table 3 (Cobos et al. 2016: 373) 
CommentAbundance: only known from its original description (Meiri et al. 2017). 
EtymologyThis specific epithet “cicak” is the Malay word for lizard. 
References
  • COBOS, ANTHONY; L. LEE GRISMER, PERRY L. WOOD, JR., EVAN S. H. QUAH, SHAHRUL ANUAR, MOHD ABDUL MUIN 2016. Phylogenetic relationships of geckos of the Hemiphyllodactylus harterti group, a new species from Penang Island, Peninsular Malaysia, and a likely case of true cryptic speciation. Zootaxa 4107 (3): 367-380 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • NGUYEN, TRUONG QUANG; QUYEN HANH DO, HANH THI NGO, ANH VAN PHAM,<br>CUONG THE PHAM, MINH DUC LE, & THOMAS ZIEGLER 2020. Two new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Hoa Binh Province, Vietnam. Zootaxa 4801 (3): 513–536 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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