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Hemiphyllodactylus dushanensis ZHOU & LIU, 1981

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Dushan slender gecko, Dushan Gypsy Gecko, Dushan Dwarf Gecko
G: Dushan-Halbblattfingergecko
Chinese: 独山半叶趾虎 
SynonymHemiphyllodactylus yunnanensis dushanensis ZHOU & LIU 1981
Hemiphyllodactylus yunnanensis — ZUG 2010 (part.)
Hemiphyllodactylus dushanensis — GRISMER et al. 2013
Hemiphyllodactylus dushanensis — GUO et al. 2015
Hemiphyllodactylus dushanensis — SUKPRASERT et al. 2018
Hemiphyllodactylus dushanensis — SUNG et al. 2018 
DistributionChina (Guizhou, Fujian)

Type locality: Dushan Junior High School in Dushan County, Guizhou Province (elevation 970 m) (China) (original in Chinese)].  
Reproductionoviparous 
TypesHolotype: NRIBAS ? 78999, male (Nanjing Normal College) 
DiagnosisDiagnosis: (dushanensis): Numbers of scansor pairs on dilated portions of digits II-V usually 3-4-4-4 in hand, 4-5-5-5 in foot*2, fifth pairs on digits III-V of foot relatively large compared to H. y. jinpingensis, extending to outer margin of digits in most specimens (Fig. 6); diameter of ear opening 0.3-0.6 mm, about 13-25% of eye diameter; dorsal surface without pattern, or with only a few irregular dark markings (from Zhou et al in Ota 1996). 
CommentSynonymy: after ZUG 2010 synonymized longlingensis, jinpingensis, and dushanensis with yunnanensis but GRISMER et al. 2013 elevated longlingensis (5 specimens), jinpingensis (4 specimens), and dushanensis (2 specimens) to full species status based on DNA sequence data but without any morphological diagnoses. Sung et al. 2018: 369 (Table 5) provide a character table for these 4 species but the characters are largely overlapping.

Distribution: see Figure 2 in GRISMER et al. 2013 for a map of this and other Hemiphyllodactylus species.

Behavior: nocturnal 
EtymologyNamed after the type locality. 
References
  • Barbour, T. 1924. A Yunnan gecko. Occasional Papers of the Boston Society of Natural History, 5:133-135 - get paper here
  • Boulenger, G.A. 1903. Descriptions of new lizards in the collection of the British Museum. Ann. Mag. Nat. Hist. (7) 12: 429-435 - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • DENG Xuejian; YE Yiyunl; YANG Daode 1998. TWO REPTILE RECORDS NEW TO HUNAN PROVINCE. Sichuan Journal of Zoology 17(2): 862 - get paper here
  • GRISMER, L. LEE; AWAL RIYANTO, DJOKO T. ISKANDAR & JIMMY A. MCGUIRE 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Pulau Enggano, southwestern Sumatra, Indonesia. Zootaxa 3821 (4): 485–495 - get paper here
  • Grismer, L. Lee; Perry L. Wood Jr, Shahrul Anuar, Evan S. H. Quah, Mohd Abdul Muin, Chan Kin Onn, Alexandra X. Sumarli and Ariel I. Loredo 2015. Repeated evolution of sympatric, palaeoendemic species in closely related, co-distributed lineages of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) across a sky-island archipelago in Peninsular Malaysia. Zoological Journal of the Linnean Society 174 (4): 859–876; DOI: 10.1111/zoj.12254 - get paper here
  • Grismer, L. Lee; Thy Neang, Thou Chav, Perry L. Wood, Jr., Jamie R. Oaks, Jeremy Holden, Jesse L. Grismer, Thomas R. Szutz and Timothy M. Youmans 2008. Additional amphibians and reptiles from the Phnom Samkos Wildlife Sanctuary in Northwestern Cardamom Mountains, Cambodia, with comments on their taxonomy and the discovery of three new species. Raffles Bulletin of Zoology 56 (1): 161-175 - get paper here
  • Grismer, L.L., Neang, T., Chav, T. & Grismer, J.L. 2008. Checklist of the amphibians and reptiles of the Cardamom region of Southwestern Cambodia. Cambodian Journal of Natural History 2008(1): 12–28 - get paper here
  • Grismer, L.L.; PERRY L. WOOD Jr, SHAHRUL ANUAR, MOHD ABDUL MUIN, EVAN S. H. QUAH, JIMMY A. MCGUIRE,, RAFE M. BROWN,0, NGO VAN TRI and PHAM HONG THAI 2013. Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia. Zoological Journal of the Linnean Society 169: 849–880 - get paper here
  • GUO, WEIBO; KAIYA ZHOU, JIE YAN & PENG LI 2015. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from western Yunnan, China. Zootaxa 3974 (3): 377–390 - get paper here
  • NGO VAN TRI; L. LEE GRISMER, PHAM HONG THAI & P. L. WOOD, JR. 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Ba Na–Nui Chua Nature Reserve, Central Vietnam. Zootaxa 3760 (4): 539–552 - get paper here
  • NGUYEN, TRUONG QUANG; QUYEN HANH DO, HANH THI NGO, ANH VAN PHAM,<br>CUONG THE PHAM, MINH DUC LE, & THOMAS ZIEGLER 2020. Two new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Hoa Binh Province, Vietnam. Zootaxa 4801 (3): 513–536 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • SHI Lei; LIN Xuan-Long; ZHAO Hui 2011. New Record of Hemiphyllodactylus yunnanensis to Xizang Autonomous Region,China. Chinese Journal of Zoology 46 (1): 131 - get paper here
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • SUKPRASERT, APISAK; SURAPOP SUTTHIWISES, VIRAYUTH LAUHACHINDA, WUT TAKSINTUM 2018. Two new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from Thailand. Zootaxa 4369 (3): 363–376 - get paper here
  • SUNG, YIK-HEI; WING-HO LEE, HO-NAM NG, YANJIE ZHANG, JIAN-HUAN YANG 2018. A new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Hong Kong. Zootaxa 4392 (2): 361–373 - get paper here
  • Taylor, E.H. 1963. The lizards of Thailand. Univ. Kansas Sci. Bull. 44: 687-1077. - get paper here
  • Wang, C. G. M. Z. X., Jia, S. Y. K. T. L., & Hu, C. Z. J. 2022. An updated species checklist of amphibians and reptiles in Fujian Province, China. Biodiversity Science, 22090 - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Wu, Jianguo 2015. Can changes in the distribution of lizard species over the past 50 years be attributed to climate change? Theoretical and Applied Climatology, DOI<br>10.1007/s00704-015-1553-4 - get paper here
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
  • Zhou & LIU in: Zhou, KAI-YA, YUE-ZHEN LIU, and GUANG-PING YANG 1981. Three new subspecies of Hemiphyllodactylus yunnanensis (Boulenger) from China [in Chinese]. Acta Zootaxon. Sinica, Beijing, 6 (2): 202-209 [English trans in Smithsonian Herp. Inf. Service 110]. - get paper here
  • Zhou, K.-y.;Liu, Y.-z.;Yang, G-p. 1996. Three New Subspecies of Hemiphyllodactylus yunnanensis (Boulenger) from China (Lacertiformes: Gekkonidae) [English translation from Chinese by H. Ota]. Smithsonian Herp. Inf. Serv. (110) - get paper here
 
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