Hemiphyllodactylus margarethae BRONGERSMA, 1931
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Higher Taxa | Gekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) |
Subspecies | |
Common Names | E: Sumatran slender gecko |
Synonym | Hemiphyllodactylus margarethae BRONGERSMA 1931: 11 Hemiphyllodactylus margarethae — ZUG 2010: 47 |
Distribution | Indonesia (Sumatra in text; Borneo on map [Fig. 20 in ZUG 2010]). Type locality: “Fort de Kock, Sumatra” (Bukittinggi, Sumatera Barat) |
Reproduction | oviparous (manual imputation, fide Zimin et al. 2022) |
Types | Holotype: ZMA.RENA 11095 |
Diagnosis | Description. A bisexual taxon of geckos (Gekkoninae) with robust habitus, slightly compressed trunk and moderately large head (see Figures 3, 23), tail round in cross section and subequal to SVL. Adults not dimorphic: 36.0–46.9 mm SVL (mean ± SD, 40.8 mm ± 3.6], n = 8), 14.5–25.4 mm TrunkL (20.6 mm ± 3.3), 8.2–10.4 mm HeadL (9.6 mm ± 0.87), 5.8–8.2 mm HeadW (6.8 mm ± 0.86), 3.3–4.7 mm SnEye (4.1 mm ± 0.56), 2.7–3.6 mm NarEye (3.2 mm ± 0.40), 1.6–2.5 mm EyeD (2.1 mm ± 0.35), 1.4–1.9 mm SnW (1.7 mm ± 0.23). Adult proportions: 40–54% TrunkL/SVL (mean ± SD, 50.3% ± 4.8), 21–26% HeadL/SVL (23.5% ± 1.6), 15–19% HeadW/ SVL (16.7% ± 1.3), 66–79% HeadW/HeadL (71.2% ± 4.4), 39–46% SnEye/HeadL (42.6% ± 2.4), 29–35% NarEye/HeadL (33.0% ± 2.0), 16–25% EyeD/HeadL (22.0% ± 3.6), 15–19% SnW/HeadL (17.3% ± 1.4), 47–81% EyeD/NarEye (67.0% ± 12.6), 22–28% SnW/HeadW (24.3% ± 1.9%). Scalation is predominantly granular from head onto tail, both dorsally and ventrally; ventral trunk scales slightly larger than dorsal ones, 11–17 Dorsal (median ± SD, 12.5 ± 2.3) and 6–12 Ventral (7.5 ± 2.3); similarly, subcaudal scales slightly larger than dorsal caudal scales but not platelike. Cloacal spurs present, modest sized, 1–2 CloacS (2 ± 0.5). Larger scales on lips and snout, rostral largest, rectangular to pentagonal, often slightly concave on dorsomedial edge with slight cleft; 2–3 CircNa (3 ± 0.4), 2–4 SnS (3 ± 0.6); labial scales enlarged from rostral to below eye, becoming progressively smaller in subocular rictus, 10–13 Suplab (11 ± 1.2), 9–12 Inflab (10 ± 0.9); 6–11 Chin (7.5 ± 2.0), those behind mental distinctly enlarged; ear opening distinct with no bordering enlarged scales. Each digit with expanded pad, terminal two phalanges free, arising from within pad on second to fifth digits of foreand hindfoot and each clawed; pads of these digits each with large triangular apical lamella bordered proximally by lyre-shaped lamellae (scansors); modal digital formulae likely 4-4-4-4 (forefoot) and 4-5-5-5 (hindfoot) for scansors; first digit of foreand hindfeet compressed, usually 5 rectangular lamellae (4–8 fore, 4–7 hind) ventrally, terminal phalanx not free with or without minute claw. Adult females often with precloacal pores series (0–12 PreclPor), males (only one adult in sample) always with precloacal pores (median 11) always separated from femoral pore series, 0–29 (female) 26 (male) TotPore. (Zug 2010) Coloration: no images of living specimens and preserved specimens pattern indistinct owing to fading. (Zug 2010) Major diagnostic features: bisexual taxon; unpigmented caecum, oviducts pigmented or not; if present, femoral pore series separate from precloacal pore series; chin scales bordering mental and first infralabial distinctly enlarged; digital lamellae formulae 4-4-4-4 (forefoot) and 4-5-5-5 (hindfoot); average adult SVL ~41 mm. (Zug 2010) Description of holotype: An adult male (Figure 23), 38.8 mm SVL, 40 TailL, 18.7 mm TrunkL, 9.9 mm HeadL, 7.5 mm HeadW, 4.3 mm SnEye, 3.4 mm NarEye, 1.6 mm EyeD, and 1.9 mm SnW. Proportions: 48% TrunkL/SVL, 25% HeadL/SVL, 19% HeadW/SVL, 76% HeadW/HeadL, 43% SnEye/HeadL, 34% NarEye/HeadL, 16% EyeD/HeadL, 19% SnW/HeadL, 47% EyeD/NarEye, 25% SnW/HeadW. Scalation: 3 CircNa, 3 SnS, 10 Suplab, 11 Inflab, 6 Chin (anteromedial ones distinctly enlarged, see Figure 7C), 13 Dorsal, 7 Ventral, 2 CloacS, Subcaud not enlarged, 11 PreclPor, 26 TotPore, separate precloacal and femoral series, digital formulae 4-5-5-5 (forefoot) and 5-5-6-5 (hindfoot). Caecum and testis epididymis presumably unpigmented. Body ground color faded to pinkish beige, small paired dark blotches dorsally on trunk; paratype (ZMA 11096) has a hint of a postsacral mark. (Zug 2010) |
Comment | Synonymy: Has been previously synonymized with Hemiphyllodactylus typus (by BRONGERSMA 1932 and SMITH 1935) but revalidated by ZUG 2010. Behavior: nocturnal |
Etymology | The origin of the name margarethae is unknown (ZUG 2010). |
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