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Hemirhagerrhis kelleri BOETTGER, 1893

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Higher TaxaPsammophiidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Stripe-bellied Bark Snake, Keller's Bark Snake, Striped Bark Snake 
SynonymHemirhagerrhis kelleri BOETTGER 1893: 129
Hemirhagerrhis kelleri — STEJNEGER 1893: 729
Amplorhinus taeniatus STERNFELD 1908: 244
Hemirrhagerrhis [sic] kelleri — CALABRESI 1927: 54
Hemirrhaggeris [sic] kelleri — SCORTECCI 1929: 272
Hemirhargerris [sic] kelleri — UTHMÖLLER 1942: 40
Hemirhagerrhis kelleri — GANS et al. 1965: 66
Hemirhagerrhis kelleri — BROADLEY 2000
Hemirhagerrhis kelleri — LARGEN & SPAWLS 2010: 545
Hemirhagerrhis kelleri — WALLACH et al. 2014: 326
Hemirhagerrhis kelleri — SPAWLS et al. 2018: 427 
DistributionRepublic of South Sudan (RSS), Ethiopia, Somalia, Kenya, N Tanzania

Type locality: Abdallah = Abdulla, north of Webi Shebeli, Ethiopia.  
Reproductionoviparous 
TypesHolotype: SMF 19906 
DiagnosisDiagnosis (genus): “Maxillary short, bearing 8-10 teeth, which increase only slightly in size posteriorly and are followed, after a short diastema, by two slightly enlarged fangs (Bogert, 1940, Fig. 15E); teeth on dentary small and subequal. Head depressed and distinct from the neck; eye small, with a round pupil (not vertically elliptic as stated in Boettger's original diagnosis and by several subsequent authors); elongate nasal shield semidivided by an oblique cleft extending from the nostril, which is pierced vertically or dorsolaterally. Body cylindrical; scales smooth with single apical pits in 17-17-13 rows; ventrals rounded; anal divided; tail moderate to long; subcaudals paired. Hemipenes simple, papilliform, with undivided sulcus. Oviparous.” (Broadley 2000: 5)


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CommentSynonymy: BOGERT (1940) considered H. kelleri as congeneric with H. nototaenia.

Type species: Hemirhagerrhis kelleri BOETTGER 1893: 129 is the type species of the genus Hemirhagerrhis BOETTGER 1893.

Habitat: partly arboreal (Harrington et al. 2018).

Key: Broadley 2000: 13. 
EtymologyNamed after Aleksandr N. Kaznakov, a Russian naturalist who became Director of the Caucasus Museum. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Blackburn, D.G. 1993. STANDARDIZED CRITERIA FOR THE RECOGNITION OF REPRODUCTIVE MODES IN SQUAMATE REPTILES. Herpetologica 49 (1): 118-132 - get paper here
  • Boettger, O. 1893. Uebersicht der von Prof. C. Keller anlässlich der Ruspolischen Expedition nach den Somaliländern gesammelten Reptilien und Batrachier. (Schluss). Zool. Anz. 16 (417): 129- 132. - get paper here
  • Bogert, Charles M. 1942. Snakes secured by the Snyder East African Expedition in Kenya Colony and Tanganyika Territory. American Museum Novitates (1178): 1-5 - get paper here
  • Bogert,C.M. 1940. Herpetological results of the Vernay Angola Expedition. I. Snakes, including an arrangement of the African Colubridae. Bull. Amer. Mus. Nat. Hist. 77: 1-107 - get paper here
  • Boulenger, G.A. 1896. Catalogue of the snakes in the British Museum, Vol. 3. London (Taylor & Francis), xiv + 727 pp. - get paper here
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Broadley, D.G. 2000. A review of Hemirhagerrhis viperina (Bocage) (Serpentes: Colubridae), a rupicolous Psammophine snake. Madoqua 19 (2): 161-169 [1997] - get paper here
  • Broadley, D.G. & Hughes,B. 2000. A revision of the African genus Hemirhagerrhis BOETTGER 1893 (Serpentes: Colubridae). Syntarsus 6: 1-17
  • Calabresi, Enrica 1927. Anfibi e rettili raccolti nella Somalia dai Prof f. G. Stefanini e N. Puccioni (Gennaio-Luglio 1924). Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 66:14—60. - get paper here
  • Gans, C.; LAURENT, R.F. & PANDIT, H. 1965. Notes on a herpetological collection from the Somali Republic. Ann. Mus. Roy. Afr. Centr., Sér. 8vo, Tervuren, (80) Zool., (134): 1-93
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Hughes, B. 1983. African snake faunas. Bonner Zoologische Beiträge 34: 311-356 - get paper here
  • HUGHES, B. 2018. THE SNAKES OF DODOMA. African Herp News (67): 8 - get paper here
  • Lanza, B. 1990. Amphibians and reptiles of the Somali Democratic Republic: check list and biogeography. Biogeographia, 14: 407-465 [1988] - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • Largen,M.J. & Rasmussen,J.B. 1993. Catalogue of the snakes of Ethiopia (Reptilia Serpentes), including identification keys. Tropical Zoology 6: 313-434 - get paper here
  • Loveridge, Arthur 1929. East African reptiles and amphibians in the United States National Museum. Bull. US Natl. Mus. (151): 1-135 - get paper here
  • Meirte, D. 1992. Clés de détermination des serpents d'Afrique. Museum Royal d'Afrique Centrale, Tervuren Belgique Annual Series Octavo Science Zoologique 267:1-152
  • Scortecci ,G. 1939. Spedizione zoologica del Marchese Saverio patrizi nel Basso Giuba e nell'Oltre Giuba. Giugno-agosto 1934. XII. Rettili Ofidi. Annali Mus. civ. Stor. nat. Giacomo Doria 58: 263-291 - get paper here
  • Scortecci, G. 1929. Primo contributo alla conoscenza dei Rettili e degli Anfibi della Somalia italiana. Atti Soc. ital. Sci. nat. 68: 245-279
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Stejneger, L. 1893. On some collections of reptiles and batrachians from East Africa and the adjacent islands, recently received from Dr. W. L. Abbott and Mr. William Astor Chanler, with descriptions of new species. Proc. US Natl. Mus. 16: 711-741 - get paper here
  • Vats, Rajeev; Ignas Safari 2014. Diversity of Snakes at the University of Dodoma Campus, Tanzania. American Journal of Zoological Research, 2 (3): 41-45 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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