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Herpetoreas sieboldii GÜNTHER, 1860

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Siebold’s Keelback, Sikkim Keelback 
SynonymHerpetoreas sieboldii GÜNTHER 1860: 156
Herpetoreas sieboldii — GÜNTHER 1864: 257
Herpetoreas sieboldii — THEOBALD 1868: 48
Herpetoreas sieboldii — THEOBALD 1876: 172
Amphiesma sieboldii — DAS 1996: 53
Amphiesma sieboldii — DAVID et al. 2005
Amphiesma sieboldii — WALLACH et al. 2014: 33
Herpetoreas sieboldii — GUO et al. 2014
Herpetoreas sieboldii — LALREMSANGA et al. 2022 
DistributionPakistan, India (Uttar Pradesh, Sikkim, Punjab), Nepal, Myanmar (= Burma), Bangladesh, Bhutan

Type locality: "Sikkim, Himalaya (7500 feet above the level  
Reproductionoviparous 
TypesHolotype: BMNH 1946.1.13.16, a 917 mm male (Hermann, Adolphe and R. von Schlagintweit, 1854-1858). 
DiagnosisDiagnosis (genus). Semi-aquatic snakes with keeled dorsal scales. The posterior maxillary tooth longest, in a continuous series with the anterior ones. Body and tail slender, compressed. Two nasals, 1 loreal, 1 anterior, 2 posterior oculars. Scales moderately elongate, keeled, in 19 rows. Eye is moderate size (Günther, 1860) [from Guo et al. 2014]. 
CommentSynonymy: Has been synonymized with A. platyceps. Both species are very similar and characters are broadly overlapping (see also TILLACK 2003).

Type species: Herpetoreas sieboldii GÜNTHER 1860: 156 is the type species of the genus Herpetoreas GÜNTHER 1860.

For a comparison of characters across members of Herpetoreas see Das et al. 2020: 313 (Table 3) and Lalremsanga et al. 2022 (Table 2).

Key to the species of Herpetoreas: Lalremsanga et al. 2022, Nguyen et al. 2024: 179. 
EtymologyNamed after “Prof. von Siebold of Munich”. 
References
  • Basfore, B., Kalita, M. J., Sharma, N., & Boro, A. R. 2024. An updated checklist of snakes (Reptilia: Squamata) in northeastern India derived from a review of recent literature. Journal of Threatened Taxa, 16(11), 26131-26149 - get paper here
  • Das, A., Gower, D. J., & Deepak, V. 2020. Lost and found: rediscovery and systematics of the Northeast Indian snake Hebius pealii (Sclater, 1891). Vertebrate Zoology 70 (3): 305–318 - get paper here
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • David, P.; Vogel, G. & Pauwels, O.S.G. 2005. On the occurrence of Amphiesma bitaeniatum (WALL, 1925) in Vietnam, with preliminary remarks on the group of Amphiesma parallelum (BOULENGER, 1890) (Serpentes, Colubridae, Natriciniae). Salamandra 41 (4): 167-178 - get paper here
  • DAVID, PATRICK; ISHAN AGARWAL, RAMANA ATHREYA, ROSAMMA MATHEW,<br>GERNOT VOGEL & VIRAL K. MISTRY 2015. Revalidation of Natrix clerki Wall, 1925, an overlooked species in the genus Amphiesma Duméril, Bibron & Duméril, 1854 (Squamata: Natricidae). Zootaxa 3919 (2): 375–395 - get paper here
  • Günther, A. 1860. Contributions to a knowledge of the reptiles of the Himalaya mountains. - I. Descriptions of the new species. II. List of Himalayan reptiles, with remarks on their horizontal distribution. Proc. Zool. Soc. London 1860: 148-175 - get paper here
  • Günther, A. 1860. Reptiles of Siam. Proc. Zool. Soc. London 1860: 113-117 - get paper here
  • Günther, A. 1864. The Reptiles of British India. London (Taylor & Francis), xxvii + 452 pp. - get paper here
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Khan, M.S. 2002. Die Schlangen Pakistans [English edition as “A Guide to the snakes of Pakistan”]. Edition Chimaira (Frankfurt am Main), 265 pp. [review in HR 34: 400, Russ J Herp 12: 79] - get paper here
  • Kramer, E. 1977. Zur Schlangenfauna Nepals. Revue Suisse de Zoologie 84 (3): 721-761. - get paper here
  • Lalremsanga, H. T., Bal, A. K., Vogel, G., & Biakzuala, L. 2022. Molecular phylogenetic analyses of lesser known colubrid snakes reveal a new species of Herpetoreas (Squamata: Colubridae: Natricinae), and new insights into the systematics of Gongylosoma scriptum and its allies from northeastern India. SALAMANDRA, 58(2): 101-115 - get paper here
  • Liu, S., Hou, M., Zhou, H., Zuo, C., Yin, F., & Rao, D. 2024. A New Species of Herpetoreas Günther, 1860 (Serpentes: Natricidae) from Yunnan, China. Diversity, 16(12), 768 - get paper here
  • Malnate, E. V. 1966. Amphiesma platyceps (BLYTH) and Amphiesma sieboldii (GÜNTHER): sibling species (Reptilia: Serpentes). J. Bombay Nat. Hist. Soc. 63 (1): 1-17 - get paper here
  • Nguyen T.V., Lalremsanga H.T., Biakzuala L. & Vogel G. 2024. Taxonomic reassessment of the Herpetoreas xenura (Wall, 1907)(Squamata: Serpentes: Natricidae) from Myanmar with description of a new species. European Journal of Taxonomy, 932, 158-203 - get paper here
  • Theobald, WILLIAM 1868. Catalogue of reptiles in the Museum of the Asiatic Society of Bengal. J. Asiatic Soc. Bengal, Calcutta, 37 (extra number 146): (2), vi, 7-88 - get paper here
  • Theobald,W. 1876. Descriptive catalogue of the reptiles of British India. Thacker, Spink & Co., Calcutta: xiii + 238 pp. - get paper here
  • Tillack, F. 2003. Über die Verbreitung und Biologie der Himalaya-Gebirgswassernatter Amphiesma platyceps (BLYTH 1854) und einen Fall von Amphigonia retardata (Serpentes: Colubridae: Natricinae). Sauria 25 (1): 21-27 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wangyal, Jigme Tshelthrim 2019. The status of herpetofauna of Bhutan. District Forest Office, District Administration, Trashigang, Bhutan, 20-39 - get paper here
  • Ziegler, T., and Quyet, L.K. 2006. A new natricine snake of the genus Amphiesma (Squamata: Colubridae: Natricinae) from the central Truong Son, Vietnam. Zootaxa 1225: 39-56 - get paper here
 
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