Holodactylus africanus BOETTGER, 1893

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Higher Taxa	Eublepharidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies
Common Names	E: Boettger’s strong-clawed gecko, African Whole-toed Gecko G: Afrikanischer Krallengecko, Ganzfinger-Lidgecko
Synonym	Holodactylus africanus BOETTGER 1893 Holodactylus africanus — BULENGER 1895: 166 Holodactylus aculeatus CALABRESI 1915: 238 Holodactylus africanus — KLUGE 1993 Hoplodactylus africanus — RÖSLER 1995: 66 (in error) Holodactylus africanus — BROADLEY & HOWELL 1991: 8 Holodactylus africanus — RÖSLER 2000: 89 Holodactylus africanus — ERNST in SEUFER et al. 2005 Holodactylus africanus — SPAWLS et al. 2018: 70
Distribution	S/E Ethiopia, Kenya, Somalia, NE Tanzania (Webi Shebeli/Abdulla) Type locality: Abdulla, Somalia
Reproduction	oviparous
Types	Holotype: SMF 59442
Diagnosis	Diagnosis (genus). Digits free, long, slender, compressed, not dilated, tips somewhat tapering, covered above with scales, not denticulate laterally, below granular, clawed, the claws between four scales, claws of fingers very long and strong. Pupil vertical; upper and lower eyelids well developed, connivent; dorsal lepidosis of small unequal or subequal, smooth, juxtaposed granules; tail swollen but depressed, constricted anteriorly, tapering posteriorly, covered above and below with small, juxtaposed, granular scales. Males without preanal or femoral pores [Loveridge 1947: 21]
Comment	Type species: Holodactylus africanus BOETTGER 1893 is the type species of the genus Holodactylus BOETTGER 1893.
Etymology	Named after its African origin.
References	Boettger,O. 1893. Übersicht der von Prof. C. Keller anlässlich der Ruspoli'schen Expedition nach den Somaliländern gesammelten Reptilien und Batrachier. Zool. Anz. 16 (416): 113-119 - get paper here Boulenger, G. A. 1895. On the reptiles and batrachians obtained by Mr. E. Lort-Phillips in Somaliland. Ann. Mag. nat. Hist. (6) 16: 165-169 - get paper here Boulenger, G.A. 1896. A list of the reptiles and batrachians collected by the late Prince Eugenio Ruspoli in Somaliland and Gallaland in 1893. Annali Mus. civ. Stor. nat. Genova, Giacomo Doria, (2) 17: 5-14 [11] - get paper here Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70 Calabresi, Enrica 1915. Contributo alla conoscenza dei rettili della Somalia. Monitore Zoologico Italiano (supplemento), 28:234—247. - get paper here Drewes, R.C. 1971. Notes on the Distribution of Holodactylus africanus. Jour. East Afr. Nat. Hist. Soc. 126 (28): 1-3 Grismer,L.L. 1997. Eublepharid Geckos - Living Relics of Gekkotan Evolution. Fauna 1 (1): 26-33 Kirchhof S, Wasonga V, Mazuch T, Spawls S, Malonza KP. 2023. An annotated checklist of the herpetofauna of the Sibiloi National Park in northern Kenya based on field surveys. Amphibian & Reptile Conservation 17(1/2) [General Section]: 1–18 (e324) - get paper here Lanza, B. 1990. Amphibians and reptiles of the Somali Democratic Republic: check list and biogeography. Biogeographia, 14: 407-465 [1988] - get paper here Largen, M.J.; Spawls, S. 2006. Lizards of Ethiopia (Reptilia Sauria): an annotated checklist, bibliography, gazetteer and identification. Tropical Zoology 19 (1): 21-109 - get paper here Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp. Loveridge, A. 1947. Revision of the African lizards of the family Gekkondiae. Bull. Mus. Comp. Zool. Harvard 98: 1-469 - get paper here Mazuch, Tomáš 2013. Amphibians and Reptiles of Somaliland and Eastern Ethiopia. Tomáš Mazuch Publishing, 80 pp. ISBN: 978-80-905439-0-4 - get paper here Mertens, R. 1967. Die herpetologische Sektion des Natur-Museums und Forschungs-Institutes Senckenberg in Frankfurt am Main nebst einem Verzeichnis ihrer Typen. Senckenbergiana Biologica 48: 1-106 - get paper here Parker, H. W. 1942. The lizards of British Somaliland. Bull. Mus. Comp. Zool. Harvard 91: 1—101 - get paper here Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153 Rösler, Herbert 1995. Geckos der Welt - Alle Gattungen. Urania, Leipzig, 256 pp. Scortecci, G. 1930. Secondo contributo alla conoscenza dei rettili della Somalia Italiana. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 69:127—152. [1931] - get paper here Seufer, H.; Y. Kaverkin & A. Kirschner (eds.) 2005. Die Lidgeckos. Kirschner und Seufer Verlag, 238 pp. Spawls, S. & Rotich, D. 1997. An annotated checklist of the lizards of Kenya. J. East African Nat. Hist. 86: 61-83 Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here Spawls, Stephen; Tomáš Mazuch& Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
External links	IUCN Red List National Center for Biotechnology Information http://www.gekkota.com/Photos/photos.html http://www.biodiversityexplorer.org/reptiles/squamata/agamidae/Agama_atra… http://www.geocities.com/Heartland/Prairie/3470/HOLODACTYLUS.html http://www.seh-herpetology.org/sites/seh-herpetology.org/files/uploads… Google images

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