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Homalophis doriae PETERS, 1871

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Higher TaxaHomalopsidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Sarawak Mud Snake, Sarawak Water Snake, Blotched-Lipped Mud Snake 
SynonymHomalophis Doriae PETERS 1871: 577
Hypsirhina doriae — GYI 1970
Enhydris doriae — BAUER et al. 1995: 72
Enhydris cf. doriae — AULIYA 2006
Enhydris doriae — MURPHY 2007: 106
Enhydris doriae — WALLACH et al. 2014: 270
Homalophis doriae — MURPHY & VORIS 2014: 23
Homalophis doriae — BERNSTEIN et al. 2023 
DistributionMalaysia (Borneo: Sarawak)

Type locality: “Sarawak”  
Reproductionovovivparous 
TypesSyntypes: ZMB 7120, MSNG 30665 
DiagnosisDiagnosis (genus): Large bodied, smooth scales; nasal scales in contact; 25–33 rows of dorsal scales, subocular scales, divided internasal; first labial does not contact loreal; 2–4 pairs of chin shields, 2–3 supraoculars; 2–3 postoculars; 13–15 upper labials, upper labials behind eye are horizontally divided into three tiers; lower labials 15–20, dorsum uniform brown or gray with red or yellow ventral surface (MURPHY & VORIS 2014). 
CommentType species: Homalophis Doriae PETERS 1871: 577 is the type species of the genus Homalophis PETERS 1871.

Habitat: aquatic; swamps, muddy rivers, rice fields, ponds. 
EtymologyNamed after Marquis Doria, a Genoese nobleman who supported numerous collecting expeditions into Southeast Asia and who collected the syntypes of this species. 
References
  • Auliya, M. 2006. Taxonomy, Life History, and conservation of giant reptiles in west Kalimantan. Natur und Tier Verlag, Münster, 432 pp. - get paper here
  • Bauer, A.M.; Günther,R. & Klipfel,M. 1995. The herpetological contributions of Wilhelm C.H. Peters (1815-1883). SSAR Facsimile Reprints in Herpetology, 714 pp.
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bernstein, J. M., de Souza, H. F., Murphy, J. C., Voris, H. K., Brown, R. M., Myers, E. A., Harrington, S., Shanker, K., & Ruane, S. 2023. Phylogenomics of Fresh and Formalin Specimens Resolves the Systematics of Old World Mud Snakes (Serpentes: Homalopsidae) and Expands Biogeographic Inference. Bulletin of the Society of Systematic Biology - get paper here
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • Gyi, Ko Ko 1970. A revision of colubrid snakes of the sub-family Homalopsinae. Univ. Kans. Publs. Mus. Nat. Hist 20 (2): 44-223 - get paper here
  • Murphy, J.C. & Voris, H.K. 2014. A Checklist and Key to the Homalopsid Snakes (Reptilia, Squamata, Serpentes), with the Description of New Genera. FIELDIANA: LIFE AND EARTH SCIENCES (8): 1–43 - get paper here
  • Murphy, John C. 2007. Homalopsid Snakes: Evolution in the Mud. Krieger Publishing, Malabar, Florida, 249 pp.
  • Peters, W. 1871. Uber neue Reptilien aus Ostafrika und Sarawak (Borneo), vorzüglich aus der Sammlung des Hrn. Marquis J. Doria zu Genua. Mber. k. preuss. Akad. Wiss., Berlin [1871]: 566-581 - get paper here
  • Stuebing, R.B., Inger, R.F. & Lardner, B. 2014. A field guide to the snakes of Borneo, second edition. Natural history Publications (Borneo)
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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