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Homalopsis semizonata BLYTH, 1855

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Higher TaxaHomalopsidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Martaban Water Snake 
SynonymHomalopsis semizonata BLYTH 1855: 187
Pythonia semizonata — BLYTH 1859: 279
Pythonella semizonata — CARLLEYLE, 1869: 200 (lapsus for Pythonia)
Homalopsis buccata – THEOBALD 1868: 58 (part.)
Homalopsis buccata – THEOBALD 1868: 66 (part.)
Homalopsis buccata – NICHOLSON 1874: 62 (part.)
Homalopsis buccata – THEOBALD 1876: 185 (part.)
Homalopsis buccata – BOULENGER 1887: 475 (part.)
Homalopsis buccata – SCLATER 1891: 53 (part.)
Homalopsis buccata – BOULENGER 1893: 309 (part.)
Homalopsis buccata – BOULENGER 1896: 16 (part.)
Homalopsis buccata – WALL & EVANS 1901: 616 (part.)
Homalopsis buccata – WALL 1903: 94 (part.)
Homalopsis buccata – ROOIJ 1917: 186 (part.)
Homalopsis buccata – WALL 1923: 38 (part.)
Homalopsis buccata – HUNDLEY 1964: 22 (part.)
Homalopsis buccata – DOWLING & JENNER 1988: 4 (part.)
Homalopsis buccata – MURPHY 2007: 193 (part.)
Homalopsis semizonata — MURPHY & VORIS 2014: 25
Homalopsis semizonata — WALLACH et al. 2014: 335 
DistributionMyanmar

Type Locality: Mottama (formerly Martaban), Myanmar.  
Reproductionviviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZSI 8137, collected by Major Berdmore 
DiagnosisDiagnosis: Homalopsis semizonata has a divided or fragmented loreal contacting upper labials 1–4 or 1–5; three prefrontals; 39–44 dorsal scale rows at midbody, reduced to more than 30 posteriorly; one postocular and one postsubocular. Homalopsis buccata has a single loreal contacting upper labials 1-4; two prefrontals; 33–40 dorsal scale rows at midbody, reduced to less than 30 posteriorly; H. hardwickii has two prefrontal scales; 39 scale rows at midbody reduced to 28 posteriorly. Homalopsis nigroventralis has upper labials 1–3 contacting the loreal; two pre- frontals; 35–39 dorsal scale rows at midbody, reduced to less than 30 posteriorly; one or two postocular scales plus one postsubocular scale; and a reverse color pattern on the venter (dark olive-gray with white spots). H. mereljcoxi has a single loreal contacting upper labials 1–4; two prefrontals; 40–47 scale rows at midbody, reduced to 30 or more posteriorly; two postocular scales plus a postsubocular [MURPHY et al. 2012].


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CommentSynonymy after MURPHY et al. 2012. Homalopsis mereljcoxi and H. semizonata may be synonyms of H. buccata, given their limited genetic differentiation (Bernstein et al. 2021). In fact, The former two species nest within H. buccata. 
References
  • Blyth, E. 1855. Collections presented by Capt. Beddome and Mr. Theobald. J. Asiatic Soc. Bengal, Calcutta, 24: 187 - get paper here
  • Blyth, E. 1859. Of reptiles,... J. Asiatic Society of Bengal, 28:279 - get paper here
  • Boulenger, G.A. 1887. An account of the reptiles and batrachians obtained in Tenasserim by M. L. Fea, of the Genova Civic Museum. Ann. Mus. civ. stor. nat. Genova, 2. Ser. 5: 474-486 - get paper here
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Boulenger, G.A. 1896. Catalogue of the snakes in the British Museum, Vol. 3. London (Taylor & Francis), xiv + 727 pp. - get paper here
  • Carlleyle, A.C.L. 1869. Descriptions of two new species, belonging to the genera Varanus, and Feranioides respectively, from near Agra. J. Asiat. Soc. Bengal 38: 192-200 - get paper here
  • Dowling, H.G., & Jenner, J.V. 1988. Snakes of Burma: checklist of reported species and bibliography. Smithsonian Herp. Inf. Serv. (76): 19 pp. - get paper here
  • Hundley, H.G. 1964. Checklist of Burmese Reptiles. H.G. Hundley, Kyi, Burma, 32 pp.
  • Murphy, J.C. & Voris, H.K. 2014. A Checklist and Key to the Homalopsid Snakes (Reptilia, Squamata, Serpentes), with the Description of New Genera. FIELDIANA: LIFE AND EARTH SCIENCES (8): 1–43 - get paper here
  • Murphy, John C. 2007. Homalopsid Snakes: Evolution in the Mud. Krieger Publishing, Malabar, Florida, 249 pp.
  • MURPHY, JOHN C.; HAROLD K. VORIS, B.H.C.K. MURTHY, JOSHUA TRAUB & CHRISTINA CUMBERBATCH 2012. The masked water snakes of the genus Homalopsis Kuhl & van Hasselt, 1822 (Squamata, Serpentes, Homalopsidae), with the description of a new species. Zootaxa 3208: 1–26 - get paper here
  • Sclater, W. L. 1891. List of snakes in the Indian Museum. Trustees of the Indian Museum, Calcutta, 79 pp. - get paper here
  • Theobald, William 1868. Catalogue of the reptiles of British Birma, embracing the provinces of Pegu, Martaban, and Tenasserim; with descriptions of new or little-known species. Zool. J. Linnean Soc. 10: 4-67. - get paper here
  • Theobald, WILLIAM 1868. Catalogue of reptiles in the Museum of the Asiatic Society of Bengal. J. Asiatic Soc. Bengal, Calcutta, 37 (extra number 146): (2), vi, 7-88 - get paper here
  • Theobald,W. 1876. Descriptive catalogue of the reptiles of British India. Thacker, Spink & Co., Calcutta: xiii + 238 pp. - get paper here
  • Wall, F. 1903. A prodromus of the snakes hitherto recorded from China, Japan, and the Loo Choo Islands, with some notes. Proc. Zool. Soc. London 1903: 84-102 - get paper here
  • Wall, F. 1923. How to Identify the Snakes of India. F. Wall, Karachi, 56 pp
  • Wall, F. and Evans, G.H. 1901. Notes on Ophidia collected in Burma from May to December, 1899. J. Bombay nat. Hist. Soc. 13: 343-354 [1900] - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
  • Zug, G.R. & Mulcahy, D.G. 2019. Identification guide Amphibians and reptiles of South Tanintharyi. Fauna & Flora International, 101 pp.
 
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