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Homonota horrida (BURMEISTER, 1861)

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Higher TaxaPhyllodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: South American Marked Gecko 
SynonymGymnodactylus horridus BURMEISTER 1861
Wallsaurus horridus — UNDERWOOD 1954: 475
Homonota horrida — KLUGE 1964 (partim)
Gymnodactylus (Wallsaurus) horridus — WERMUTH 1965: 54
Gymnodayctylus pasteuri — WERMUTH 1965: 63 (nomen novum)
Homonota pasteuri — WERMUTH 1965: 201 (nomen novum)
Homonota horrida — GALLARDO 1966: 16
Homonota horrida — GALLARDO 1972
Homonota horrida — PETERS & DONOSO-BARROS 1970
Homonota horrida — CEI 1993
Homonota horrida — RÖSLER 1995: 127
Homonota horrida — RÖSLER 2000: 89
Homonota horrida — RÖSLER 2017: 81
Homonota horrida — CACCIALI et al. 2017
Homonota horrida — AVILA et al. 2021 
DistributionS Argentina (Rio Negro, La Rioja), Paraguay (Chaco)

Type locality: Mendoza (“in den Schluchten der Sierra bei Challao”), Argentina.  
ReproductionOviparous. 
TypesLectotype: MLUH IZH R 1 (Fig. 9 in Cacciali et al. 2017); paralectotype IZH-R 2, designation according to Müller 1941. 
DiagnosisDiagnosis: A large species of Homonota with a dark dorsal color (grey or brown) with a pattern of clear transversal bands connected with a vertebral stripe. Additionally, it is differentiated from any other Homonota by the large size and development of the keeled scales on the head (including laterals) and dorsum [Cacciali et al. 2017: 14]. 
CommentSynonymy mainly after CEI 1993. H. horrida has been synonymized with H. fasciata (ABDALA & LAVILLA 1993, CACCIALI et al. 2007) but revalidated by
Cacciali et al. 2017 (see also H. fasciata).

Distribution: reports from Brazil (Matto Grosso) need to be revisited after the revision of Cacciali et al. 2017. Reports from SE Bolivia (Cochabamba, Santa Cruz, Tarija) are probably H. septentrionalis.

Type species: Gymnodactylus horridus BURMEISTER 1861 is the type species of the genus Wallsaurus UNDERWOOD 1954. 
References
  • Abdala V. Lavilla E O. 1994. Synonymy and chresonymy of Homonota fasciata (Sauria: Gekkonidae) [in Spanish]. Acta Zoologica Lilloana 42 (2): 279-282
  • Abdala, V. and E. O. Lavilla. 1993. Homonota fasciata (Duméril y Bibrton, 1836), nombre válido para Homonota pasteuri Wermuth, 1965 y Homonota horrida (Burmeister, 1861) (Sauria: Gekkonidae). Acta Zoológica Lilloana 42: 279–282
  • Aun, L.;Martori, R. 1994. Biología de una poblacion de Homonota horrida. Cuadernos de Herpetología 8 (1): 90-96 - get paper here
  • AVILA, LUCIANO JAVIER; CRISTIAN HERNAN FULVIO PEREZ, IGNACIO MINOLI & MARIANA MORANDO 2012. A new species of Homonota (Reptilia: Squamata: Gekkota: Phyllodactylidae) from the Ventania mountain range, Southeastern Pampas, Buenos Aires Province, Argentina. Zootaxa 3431: 19–36 - get paper here
  • AVILA, LUCIANO JAVIER; LORENA ELIZABETH MARTINEZ & MARIANA MORANDO 2013. Checklist of lizards and amphisbaenians of Argentina: an update. Zootaxa 3616 (3): 201–238 - get paper here
  • Avila, Luciano Javier; Martín Miguel Femenías, Cristian Hernán Fulvio Pérez, Mariana Morando 2021. Nuevas especies de Liolaemus para la herpetofauna de la provincia de La Rioja, Argentina. Cuadernos de Herpetología 35 (S1): 223-228
  • Berg, C. 1895. Dos reptiles neuvos descritos. Anales del Museo Nacional de Buenos Aires, 4:189-194. - get paper here
  • Böhm, Stephan 2015. Eindrücke der Herpetofauna im Chaco Paraguays. Terraria-Elaphe 2015 (1): 40-46 [2014] - get paper here
  • Burmeister,H. 1861. Reise durch die La Plata Staaten mit besonderer Rücksicht auf die physische Beschaffenheit und den Culturzustand der Argentinischen Republik. Ausgeführt in den Jahren 1857, 1858, 1859 und 1860. Halle, H.W. Schmidt, 2: iv + 538 pp.
  • Cacciali P, Morando M, Medina CD, Köhler G, Motte M, Avila LJ. 2017. Taxonomic analysis of Paraguayan samples of Homonota fasciata Duméril & Bibron (1836) with the revalidation of Homonota horrida Burmeister (1861) (Reptilia: Squamata: Phyllodactylidae) and the description of a new species. PeerJ 5:e3523 - get paper here
  • Cacciali, P.; Ávila, I. & Bauer, F. 2007. A new species of Homonota (Squamata, Gekkonidae) from Paraguay, with a key to the genus. Phyllomedusa 6 (2): 137-146 - get paper here
  • Cacciali, Pier; Norman J. Scott, Aida Luz Aquino Ortíz, Lee A. Fitzgerald, and Paul Smith 2016. The Reptiles of Paraguay: Literature, Distribution, and an Annotated Taxonomic Checklist. SPECIAL PUBLICATION OF THE MUSEUM OF SOUTHWESTERN BIOLOGY, NUMBER 11: 1–373 - get paper here
  • CAJADE, RODRIGO; EDUARDO GABRIEL ETCHEPARE, CAMILA FALCIONE, DIEGO ANDRÉS BARRASSO, BLANCA BEATRIZ ALVAREZ 2013. A new species of Homonota (Reptilia: Squamata: Gekkota: Phyllodactylidae) endemic to the hills of Paraje Tres Cerros, Corrientes Province, Argentina. Zootaxa 3709 (2): 162–176 - get paper here
  • Cei, J. M. 1978. Homonota andicola, nueva especie de Gekkonidae (Sauria, Reptilia) de la regiôn Andina de Uspallata, Argentina. Publicaciones Ocasionales Instituto de Biologia Animal, Universidad Nacional de Cuyo, Mendoza, Argentina, (1):1—2
  • Cei, J. M. 1993. Reptiles del noroeste, nordeste y este de la Argentina. Museo Regionale Sci. Naturale Torino, Monografie 14: 1-949
  • Dirksen, L. & De la Riva, I. 1999. The lizards and amphisbaenians of Bolivia (Reptilia, Squamata): checklist, localities, and bibliography. Graellsia 55: 199-215 - get paper here
  • Duméril, A. M.C., G. BIBRON & A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 9. Paris, XX + 440 S. - get paper here
  • Duméril, A.M. C. and G. Bibron. 1836. Erpetologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 3. Libr. Encyclopédique Roret, Paris, 528 pp. - get paper here
  • Etchepare, Eduardo G.; María del Rosario Ingaramo, Camila Falcione, Roberto H. Aguirre,Carlos E. Barrios 2011. Homonota fasciata Duméril y Bibron, 1839 (Reptilia,Squamata, Phyllodactylidae). Primer registro para la provinciade Corrientes (República Argentina). Cuadernos de Herpetología 25 (1): 21-22 - get paper here
  • Gallardo, J.M. 1972. Composicion faunistica de los saurios de la Provincia de la Pampa, Republica Argentina. Neotropica 17 (52): 44-48
  • Gallardo, Jose M. 1966. Liolaemus lentus nov. Sp. (Iguanidae) de la Pampa y Algunas observaciones sobre los saurios de picha provincia Argentina y del oeste de Buenos Aires. Neotropica 12 (37): 15-29
  • Gallardo, José M. 1969. Las especies de saurios (Reptilia) de la Provincia de Santa Fe, Argentina, y consideraciones sobre su ecologia y zoogeografia. Neotropica 15 (47): 73-81
  • Gonzales, Lucindo; Arturo Muñoz y Edson Cortez 2006. PRIMER REPORTE SOBRE LA HERPETOFAUNA DE LA RESERVA NATURAL “EL CORBALÁN”, TARIJA, BOLIVIA [FIRST REPORT ON THE HERPETOFAUNA OF “EL CORBALAN” NATURAL RESERVE, TARIJA, BOLIVIA]. Kempffiana 2 (1): 72-94 - get paper here
  • Ingaramo, María del Rosario; Marangoni, Federico; Cajade, Rodrigo 2015. Herpetofauna de la Reserva Paleontológica del Arroyo Toropí, Bella Vista, Corrientes, Argentina. Cuad. herpetol. 29 (1): - get paper here
  • Jerdon,T.C. 1865. Remarks on observations contained in Dr. Günther’s work on the reptiles of British India. Ann. Mag. Nat. Hist. (3) 15: 416-418 - get paper here
  • Kacoliris F.P.; Berkunsky I. & Williams J. 2006. Herpetofauna of Impenetrable, Argentinean Great Chaco. Phyllomedusa 5 (2): 149-158 - get paper here
  • Koslowsky,J. 1895. Batracios y reptiles de Rioja y Catamarca. Rev. Mus. La Plata 6: 359-370 - get paper here
  • Langstroth, R.P. 2005. Adiciones probables y confirmadas para la saurofauna boliviana. Kempffiana 1 (1): 101-128
  • Lavilla, E. O., GONZÁLES, L. & FERNÁNDEZ, I. 1996. Herpetologia Amboroense. Informe sobre la herpetofauna del Parque Nacional Amboró y zonas aledañas. Consorcio FAN-TNC, Santa Cruz. 46 pp.
  • Morando, Mariana; Cintia D. Medina, Luciano J. Avila, Cristian H. F. Perez, Amy Buxton and Jack W. Sites Jr. 2014. Molecular phylogeny of the New World gecko genus Homonota (Squamata: Phyllodactylidae). Zoologica Scripta DOI: 10.1111/zsc.12052 - get paper here
  • Müller,L. 1941. Über die in der Sammlung des zoologischen Institutes der Universität Halle a. Saale aufbewahrten Amphibien und Reptilientypen. [type catalogue] Zeitschr. f. Naturwiss. 94: 182-205
  • Pelegrin, N., Lara-Resendiz, R. A., Leao S. M. and Sanchez, J. M. 2020. Homonota horrida (South American marked gecko). Habitat. Herpetological Review 51: 125-126. - get paper here
  • Peracca, M. G. 1897. Rettili e Anfibi. Viaggio del Dott. Alfredo Borelli nel Chaco boliviano e nella Repubblica Argentina. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino, 12 (274): 1-19. - get paper here
  • Perez, Cristian H. F., Daniel R. Perez and Luciano J. Avila. 2008. Southernmost records for Homonota fasciata (Duméril & Bibron, 1836) in northern Patagonia, Argentina. Herpetozoa 20 (3/4):182-184 [2007] - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • RÖSLER, H. 2020. Hommage an Murk van Phelsum. Sauria 42 (2): 3-27
  • Rösler, Herbert 1995. Geckos der Welt - Alle Gattungen. Urania, Leipzig, 256 pp.
  • Underwood, Garth 1954. On the classifcation and evolution of geckos. Proc. Zool. Soc. London, 124 (3): 469-492
  • Valdez, F; Alés, R G; Acosta, J C; Corrales, L; Fernandez, R; Acosta, R; Martínez, T A & Blanco, G; 2019. Homonota horrida (South American Marked Gecko) Aquatic locomotion. Herpetological Review 50 (2): 374 - get paper here
 
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