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Homopus femoralis BOULENGER, 1888

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Higher TaxaTestudinidae, Testudinoidea, Testudines (turtles)
Common NamesE: Greater dwarf tortoise, Greater padloper, Karoo Tortoise
G: Große Zwergschildkröte 
SynonymHomopus femoralis BOULENGER 1888
Homopus femoralis — BOUR 1980
Homopus femoralis — VALVERDE 2005
Homopus femoralis — TTWG 2014
Homopus femoralis — TTWG 2021 
DistributionRepublic of South Africa (E Cape Province, SW Orange Free State , SW Transvaal ?)

Type locality: "Cradock," Cape Province, Republic of South Africa.  
TypesSyntype: BMNH 1946.1.22.54 and others (formerly BMNH 1888.12.28.1) 
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bonin, F., Devaux, B. & Dupré, A. 2006. Turtles of the World. English translation by P.C.H. Pritchard. Johns Hopkins University Press, 416 pp.
  • Boulenger, G.A. 1888. Description of a new land-tortoise from South Africa, from a specimen living in the Society's Gardens. Proc. Zool. Soc. London 1888: 251. - get paper here
  • Bour, R. 1980. Essai sur la taxinomie des Testudinidae actuels (Reptilia, Chelonii). Bull. Mus. natl. Hist. nat. Paris (4) 2 (2): 541-546
  • Branch, W.R.; Rau, R. 1991. Life History Note: Homopus femoralis. J. Herp. Assoc. Africa (39): 27-27 - get paper here
  • Branch, William R. 1993. A Photographic Guide to Snakes and Other Reptiles of Southern Africa. Cape Town: Struik Publishers, 144 S.
  • Conradie, Werner; Brian Reeves , Neil Brown, & Jan A. Venter 2016. Herpetofauna of the Oviston, Commando Drift and Tsolwana nature reserves in the arid interior of the Eastern Cape Province, South Africa. Indago 32: 81-98 - get paper here
  • Duerden, J.E. 1906. South African tortoises of the genus Homopus, with description of a new species. Records of the Albany Museum 1: 405-411 - get paper here
  • Ernst,C.H. and Barbour,R.W. 1989. Turtles of the World. Smithsonian Institution Press, Washington D.C. - London
  • Greig, J.C., and P.D. Burdett. 1976. Patterns in the distributions of Southern African terrestrial tortoises (Cryptodira: Testudinidae). Zool. Africana 11(2): 250-267.
  • Hofmeyr, Margaretha D. & William R. Branch 2018. The padloper’s tortuous path (Chelonia: Testudinidae): Two genera, not one. African Journal of Herpetology 67 (2): 99-112 - get paper here
  • Hofmeyr, Margaretha D.; Melita Vamberger, William Branch, Alfred Schleicher and Savel R. Daniels 2016. Tortoise (Reptilia, Testudinidae) radiations in Southern Africa from the Eocene to the present. Zoologica Scripta 46: 389–400, DOI: 10.1111/zsc.12223 [Record online: 18 NOV 2016] - get paper here
  • KUPERUS, S., & LOEHR, V. 2009. Homopus femoralis Boulenger, 1888 - Nesting. African Herp News (49): 10-11 - get paper here
  • Lafleur, Y. et al. 1999. CITES Identification Guide: Turtles and Tortoises. ISBN 957-02-5259-6
  • Loehr, V J T; 2019. Homopus femoralis (Greater Padloper) Reproduction and growth. Herpetological Review 50 (3): 517-518 - get paper here
  • LOEHR, V. J. T. 2012. Homopus femoralis Boulenger, 1888 Diet. African Herp News (57): 9 - get paper here
  • Loehr, Victor J. T. 2012. Road Mortality in the Greater Padloper, Homopus femoralis (Testudinidae). Chelonian Conservation and Biology Dec 2012, Vol. 11, No. 2: 226-229. - get paper here
  • Loehr, Victor J. T. 2013. Homopus femoralis (greater padloper) reproduction. Herpetological Review 44 (2): 304-305 - get paper here
  • Loehr, Victor J.T. 2013. Activity of the Greater Padloper, Homopus femoralis (Testudinidae), in Relation to Rainfall. African Zoology Oct 2012, Vol. 47, No. 2: 294-300. - get paper here
  • Loveridge, Arthur & Williams, Ernest E. 1957. Revision of the African Tortoises and Turtles of the Suborder Cryptodira. Bull. Mus. Comp. Zool. Harvard 115 (6): 163-557 - get paper here
  • SALES-NUNES, PEDRO M.; ANTOINE FOUQUET, FELIPE F. CURCIO, PHILIPPE J. R. KOK and MIGUEL TREFAUT RODRIGUES 2012. Cryptic species in Iphisa elegans Gray, 1851 (Squamata: Gymnophthalmidae) revealed by hemipenial morphology and molecular data. Zoological Journal of the Linnean Society 361–376 October 2012 Volume 166, Issue 2, 361-376 - get paper here
  • TTWG [Peter Paul van Dijk, John B. Iverson, Anders G.J. Rhodin, H. Bradley Shaffer, and Roger Bour] 2014. Turtles of the World, 7th Edition: Annotated Checklist of Taxonomy, Synonymy, Distribution with Maps, and Conservation Status. 000.v7. Chelonian Research Monographs (ISSN 1088-7105) No. 5, doi:10.3854/crm.5.000.checklist.v7.2014 - get paper here
  • TTWG; Rhodin, A.G.J., Iverson, J.B., Bour, R., Fritz, U., Georges, A., Shaffer, H.B., and van Dijk, P.P. 2021. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (9th Ed.). In: Rhodin, A.G.J., Iverson, J.B., van Dijk, P.P., Stanford, C.B., Goode, E.V., Buhlmann, K.A., and Mittermeier, R.A. (Eds.). Chelonian Research Monographs 8:1–472. doi:10.3854/crm.8.checklist.atlas.v9.2021. - get paper here
  • Valverde, J. 2005. African Tortoises. Reptilia (GB) (43): 12-20 - get paper here
  • Valverde, J. 2005. Afrikanische Landschildkröten. Reptilia (Münster) 10 (56): 18-25 - get paper here
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