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Ialtris haetianus (COCHRAN, 1935)

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Higher TaxaColubridae (Dipsadinae), Alsophiini, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
SubspeciesIaltris haetianus haetianus (COCHRAN 1935)
Ialtris haetianus perfector (SCHWARTZ & THOMAS 1965)
Ialtris haetianus vaticinata (SCHWARTZ 1970) 
Common NamesE: Hispaniolan Upland Racer, Haitian Ground Snake 
SynonymDarlingtonia haetiana COCHRAN 1935: 375
Darlingtonia haetiana — SCHWARTZ & HENDERSON 1991: 598
Darlingtonia haetianus — CROTHER 1999
Ialtris haetianus — HEDGES et al. 2009
Darlingtonia haetiana — GRAZZIOTIN et al. 2012
Darlingtonia haetiana — WALLACH et al. 2014: 209
Ialtris haetianus — KRYSKO et al. 2015

Ialtris haetiana haetiana (COCHRAN 1935)
Darlingtonia haetiana haetiana — SCHWARTZ & HENDERSON 1991: 598
Ialtris haetianus haetianus — HEDGES et al. 2009

Ialtris haetianus perfector (SCHWARTZ and THOMAS 1965)
Darlingtonia haetiana perfector — SCHWARTZ & HENDERSON 1991: 598
Ialtris haetianus perfector — HEDGES et al. 2009

Ialtris haetianus vaticinata (SCHWARTZ 1970)
Darlingtonia haetiana vaticinata — SCHWARTZ & HENDERSON 1991: 599
Ialtris haetianus vaticinatus — HEDGES et al. 2009 
DistributionHispaniola (Haiti, SW Dominican Republic)

haetiana: Hispaniola: Haiti, Massif de la Hotte; elevation 1000 - 5000 ft.; Type locality: Roche Croix, northeast foothills, Massif de la Hotte ( = Pic Macaya), Département de la Grand'Anse, Haiti.

perfector: Hispaniola: Sierra de Baoruco and southern versant of Massif de la Selle; elevation 2100 - 4400 ft.

vaticinata: Hispaniola: Haiti, the Montagne Noire; elevation 5000 - 5600 ft.

TypesHolotype: MCZ 38252, collected by: Philip J. Darlington, Jr. Collection Date: October 1934
Holotype: MCZ 77217 [perfector]
Holotype: MCZ 92099 [vaticinata] 
DiagnosisDiagnosis (genus Darlingtonia): Strongly bilobed, semicalyculated and semicapitated hemipenis with the capitulum restricted to the sulcate and lateral surface of the lobes, formed by papillate calyces; relatively long lobes, representing almost half the total length of the organ [GRAZZIOTIN et al. 2012]. 
CommentThis species may be reassigned to Arrhyton (see Hedges and Garrido 1992).

Type species: Darlingtonia haetiana COCHRAN 1935 is the type species of the genus Darlingtonia COCHRAN 1935. GRAZZIOTIN et al. 2012 et al. argued that the synonymization of Darlingtonia with Ialtris (Hedges et al., 2009) is not justified given the significantly distinct, external and hemipenial morphologies of these genera. Further, Darlingtonia differs from Ialtris in having eight infralabials (nine in Ialtris), 132–144 ventrals (vs. 160– 192) and 40–54 subcaudals (vs. 57–115), an entire anal scale (vs. divided), no loreal scale (vs. present) and ungrooved teeth (vs. grooved) (data from Cochran, 1941; Schwartz and Rossman, 1976; and Schwartz and Henderson, 1991). 
EtymologyNamed after the type locality  
  • Cochran, D. M. 1935. New reptiles and amphibians collected in Haiti by P. J. Darlington. Proc. Boston Soc. Nat. Hist. 40: 367-375. - get paper here
  • Grazziotin, Felipe G.; Hussam Zaher, Robert W. Murphy, Gustavo Scrocchi, Marco A. Benavides, Ya-Ping Zhang and Sandro L. Bonatto 2012. Molecular phylogeny of the New World Dipsadidae (Serpentes: Colubroidea): a reappraisal. Cladistics DOI: 10.1111/j.1096-0031.2012.00393.x - get paper here
  • Hedges, S. B. and O. H. Garrido. 1992. Cuban snakes of the genus Arrhyton: two new species and a reconsideration of A. redimitum Cope. Herpetologica 48: 168-177 - get paper here
  • Hedges, S. Blair; Arnaud Couloux; Vidal, Nicolas 2009. Molecular phylogeny, classification, and biogeography of West Indian racer snakes of the Tribe Alsophiini (Squamata, Dipsadidae, Xenodontinae). Zootaxa 2067: 1-28 - get paper here
  • KRYSKO, KENNETH L.; DAVID W. STEADMAN, LEROY P. NUÑEZ & DAVID S. LEE 2015. Molecular phylogeny of Caribbean dipsadid (Xenodontinae: Alsophiini) snakes, including identification of the first record from the Cay Sal Bank, The Bahamas. Zootaxa 4028 (3): 441–450 - get paper here
  • Schwartz, A. 1970. Darlingtonia haetiana (Serpentes, Colubridae): a new subspecies. Herpetologica 26 (3): 324-31. - get paper here
  • Schwartz, A. 1986. Darlingtonia, D. Haitiana. Catalogue of American Amphibians and Reptiles ( 390 - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Schwartz, A. and Thomas, R. 1965. The genus Darlingtonia (Serpentes) in Hispaniola, including a new subspecies from the Dominican Republic. Breviora (229):1-10. - get paper here
  • Uetz, P.; Darko, Y.A.; Voss, O. 2023. Towards digital descriptions of all extant reptile species. Megataxa 010 (1): 027–042 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
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