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Ichnotropis chapini SCHMIDT, 1919

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Higher TaxaLacertidae, Eremiadinae, Sauria, Lacertoidea, Squamata (lizards)
Subspecies 
Common Names 
SynonymIchnotropis chapini SCHMIDT 1919
Ichnotropis chapini — WITTE 1933
Ichnotropis chapini — WITTE & LAURENT 1942: 173
Ichnotropis capensis chapini — LAURENT 1952
Ichnotropis chapini — EDWARDS et al. 2013 
DistributionNE Democratic Republic of the Congo (Zaire), Republic of South Sudan (RSS)

Type locality: Africa: Belgian Congo: Aba, Uelle Region  
Reproductionoviparous 
TypesHolotype: AMNH 10674 
DiagnosisDiagnosis: Habitus as in Ichnotropis capensis, to which it is closely allied. Hind leg not reaching axilla; head shields very rugose; frontonasd undivided; an occipital; a supraloreal between the frontonasal and the anterior loreal; an auricular shield. Anterior border of frontal square (Schmidt 1919: 508).

Description: Habitus lacertiform; body slender; legs short; tail .56 of the total length (.57 in capensis). Head shields very rugose, except nasals and anterior loreals, about as in squamulosa, more so than in maerolepidota. Rostra1 nearly as high as wide, five-sided, pointed above, narrowly separated from the Irontonasal by the superior nasals. Nostril circumscribed by three nasals, an anterosuperior, an inferior, and a very small posterior, the latter much smaller than observed in macrolepido!a and squamulosa. A smooth supraloreal between the two upper nasals, the frontonasal, prefrontal, anterior and posterior loreals. Frontal four-sided, with two longitudinal keels, anteriorly enclosed between the prefrontals. Four supraoculars; four supraciliaries, of which the anterior exceeds the other three, separated from the supraoculars by a series of granular scales. Frontoparietals slightly exceeded by the interparietal, which is bordered behind by an occipital. Parietal shields rounded behind, with three enlarged scales on each side. Temporals small, uniform, keeled. A curved auricular bordering the ear opening anteriorly; auricular larger than in squamulosn or macrolepidota examined. Labials 8-7, above and below, four anterior to the subocular (which borders the lip) on one side, five on the other.
Dorsal scales strongly keeled and imbricate, in twenty-five longitudinal and fifty-five transverse series. Ventrals smooth, imbricate, in ten fairly regular longitudinal series and thirty cross-rows to the arms. Twenty-three scales from brachial region to the gular symphysis. Scales of the preanal area small. Fold anterior to the shoulder faint (Schmidt 1919: 508).

Coloration: General color grayish brown above. A lateral white stripe originating on the subocular, outlined above and below with black, passes above the arm, but does not reach the hind leg. A very faint dorsolateral line above this (visible only when in alcohol). Between these lines on the sides is a series of transverse black spots, two or three scales wide and half a scale long, on the tips of scales; two series of similar transverse markings dorsally, one on each side of the median line, extending to the longitudinal dorsolateral stripe. Ventral scales and chin shields white, outlined with gray, the two outer rows of ventrals punctate with brown dots. Lower and upper labials mottled with light and dark. Limbs grayish brow above, light beneath (Schmidt 1919: 510). 
CommentNot listed by BROADLEY & POYNTON 1998 for the Democratic Republic of the Congo (Zaire).

Beside the type from Aba (DRC), only known from 2 other specimens collected by Schouteden in Adra (DRC) (M. van den Berg, pers. comm., 7 Oct 2017).

Abundance: only known from the type locality (Meiri et al. 2017). 
EtymologyNamed after James P. Chapin, one of the collectors. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Berg, M.P. van den 2017. An annotated bibliographic history of Ichnotropis PETERS, 1854 (Reptilia, Lacertidae) with remarks on the validity of some of the including species. [email protected] (Eidechsen Online), 2017 [4]: 60-138. - get paper here
  • Bischoff, W. 1991. Übersicht der Arten und Unterarten der Familie Lacertidae 2. Die Gattungen Eremias, Gallotia, Gastropholis, Heliobolus, Holaspis und Ichnotropis. Die Eidechse 2 (2): 14-21 - get paper here
  • Boulenger, G.A. 1921. Monograph of the Lacertidae, vol. 2. British Museum (London), 451 pp. - get paper here
  • EDWARDS, SHELLEY; WILLIAM R. BRANCH, BIEKE VANHOOYDONCK, ANTHONY HERREL, G. JOHN MEASEY, KRYSTAL A. TOLLEY 2013. Taxonomic adjustments in the systematics of the southern African lacertid lizards (Sauria: Lacertidae). Zootaxa 3669 (2): 101–114 - get paper here
  • Laurent, R. F. 1952. Batraciens et Reptiles récemment acquis par le Musée du Congo Belge. Rev. Zool. Bot. Afr. 44: 198-203.
  • Lewin, Amir; Anat Feldman, Aaron M. Bauer, Jonathan Belmaker, Donald G. Broadley, Laurent Chirio, Yuval Itescu, Matthew LeBreton, Erez Maza, Danny Meirte, Zoltán T. Nagy, Maria Novosolov, Uri Roll, Oliver Tallowin, Jean-François Trape, Enav Vidan and 2016. Patterns of species richness, endemism and environmental gradients of African reptiles. Journal of Biogeography, doi:10.1111/jbi.12848 - get paper here
  • Mayer, W. 2013. Kommentierte Lacertiden-Liste für Europa, Afrika, den Nahen Osten inklusive der Arabischen Halbinsel und Asien. [email protected] (Eidechsen Online) 2013 [7]: 81-141 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Schmidt, Karl Patterson 1919. Contributions to the Herpetology of the Belgian Congo based on the Collection of the American Congo Expedition, 1909-1915. Part I: turtles, crocodiles, lizards, and chamaeleons. Bull. Amer. Mus. Nat. Hist. 39 (2): 385-624 - get paper here
  • Witte, G. F. de 1933. Reptiles récoltés au Conge Belge par le Dr. H. Schouteden et par M. G.-F. Witte. Ann. Mus. Conge belge Zool. Ser. 1 Tome III: 53-100.
  • Witte, G. F. de 1966. Exploration du Parc National de la Garamba. Inst. Parc nation. Congo Expl. P. N. Garamba, Miss Saeg., 48 (Rept.): 1-108 - get paper here
  • WITTE, G.-F. DE & R.F. LAURENT 1942. Liste des Lacertidae du Congo Belge et description d’une espèce nouvelle. Revue de Zoologie et de Botanique Africaines, Bruxelles, 36 (2): 165-180 - get paper here
 
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