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Lampropeltis greeri WEBB, 1961

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Higher TaxaColubridae, Colubrinae, Lampropeltini, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Greer’s Kingsnake
S: Culebra Real de Greer
G: Greers Königsnatter 
SynonymLampropeltis mexicana greeri WEBB 1961
Lampropeltis mexicana greeri — GEHLBACH & BAKER 1962: 298
Lampropeltis mexicana mexicana — GEHLBACH & MCCOY 1965: 37
Lampropeltis mexicana — GARSTKA 1982: 29
Lampropeltis mexicana greeri — HILKEN & SCHLEPPER 1998
Lampropeltis mexicana greeri — SCHMIDT & KUNZ 2005: 26
Lampropeltis mexicana greeri — BRYSON et al. 2007
Lampropeltis greeri — HANSEN & SALMON 2017: 732 
DistributionMexico (Durango, Nayarit, eastward to Zacatecas, Aguascalientes, Jalisco, Sinaloa)

Type locality: "Rancho Santa Barbara (Weicher Ranch), 29 miles west-southwest of Ciudad Durango, Durango, Mexico."  
Reproductionoviparous 
TypesHolotype: MSU 190 
DiagnosisDiagnosis: A moderate-sized (maximum TL = 1,156 mm) tricolored kingsnake most similar to L. alterna and some populations of L. leonis. It differs from nearest populations of L. alterna by the presence of a golden-brown to dark brown eye color (vs. silver-gray in L. alterna), black head cap markings (mostly absent in L. alterna), the head only slightly distinct from the neck (vs. prominently so in L. alterna), the eyes only slightly protruberant (vs. prominently so in L. alterna), and lower ventral scale counts (197–204 in L. greeri vs. 211–230 in L. alterna) (Fig. 18).
Although the ranges of L. greeri and L. leonis are separated by ~270 km, the dorsal patterns of some individuals of the highly polymorphic L. leonis are similar to those seen in L. greeri. Lampropeltis greeri differs from L. leonis in lacking pronounced pattern polymorphism (with the exception of the population from Sierra del Laurel, Jalisco, where modest levels of pattern variation are evident), the snout/head color matches the body ground color (only in some individuals of L. leonis), the presence of postocular black markings (rare in L. leonis), and the lack of ontogenetic color pattern change (often pronounced in some pattern types of L. leonis) (Figs. 23, 24) [from Hansen & Salmon 2017; 732] 
CommentSynonymy: partly after  HANSEN & SALMON 2017. Gehlbach and McCoy (1965), considered L. greeri to fall within the known variation of L. m. mexicana, a position Hansen & Salmon reject.

Distribution: see map in  HANSEN & SALMON 2017: 722 (Fig. 17). Not in Nayarit fide WOOLRICH-PIÑA et al. 2016 (who only report L. mexicana). Not in Durango fide Lemos-Espinal (2018). 
EtymologyDedicated to Mr. J. Keever Greer, one of the collectors of the holotype (Webb, 1961). 
References
  • Bryson Jr., Robert W.; Jennifer Pastorini, Frank T. Burbrink and Michael R.J. Forstner 2007. A phylogeny of the Lampropeltis mexicana complex (Serpentes: Colubridae) based on mitochondrial DNA sequences suggests evidence for species-level polyphyly within Lampropeltis. Molecular Phylogenetics and Evolution 43 (2): 674-684 - get paper here
  • Garstka,W.R. 1982. Systematics of the mexicana species group of the colubrid genus Lampropeltis, with an hypothesis [of] mimicry. Breviora (466): 1-35 - get paper here
  • Gehlbach,F.R. & Baker,,J. 1962. Kingsnakes allied with Lampropletis mexicana: taxonomy and natural history. Copeia 1962 (2): 291-300 - get paper here
  • Gehlbach,F.R. & McCoy,C. 1965. Additional observations on variation and distribution of the gray-banded kingsnake, L. mexicana (Garman). Herpetologica 21 (1): 35-38 - get paper here
  • Hansen, R. W., and G. T. Salmon. 2017. Distribution analysis, taxonomic updates, and conservation status for the Lampropeltis mexicana group (Serpentes: Colubridae). Mesoamerican Herpetology 4(4): 700–758 - get paper here
  • Hansen, R.W. & Alamillo, J.A. 2018. Geographic Distribution: Lampropeltis greeri (Durango Mountain Kingsnake). Herpetological Review 49 (1): 77. - get paper here
  • Hilken,G. & Schlepper,R. 1998. Der Lampropeltis mexicana-Komplex (Serpentes: Colubridae): Naturgeschichte und Terrarienhaltung. Salamandra 34 (2): 97-124 - get paper here
  • Lemos-Espinal JA, Smith GR 2020. A checklist of the amphibians and reptiles of Sinaloa, Mexico with a conservation status summary and comparisons with neighboring states. ZooKeys 931: 85-114 - get paper here
  • Loc-Barragán JA, Smith GR, Woolrich-Piña GA, Lemos-Espinal JA 2024. An updated checklist of the amphibians and reptiles of Nayarit, Mexico with conservation status and comparison with adjoining States. Herpetozoa 37: 25-42 - get paper here
  • Schmidt, D. & Kunz, K. 2005. Ernährung von Schlangen. Natur und Tier Verlag, Münster, 159 pp. - get paper here
  • Webb,R. 1961. A new kingsnake from Mexico, with remarks on the mexicana group of the genus Lampropeltis. Copeia 1961 (3): 326-333 - get paper here
 
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