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Laudakia nupta (DE FILIPPI, 1843)

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Higher TaxaAgamidae (Agaminae), Sauria (lizards) 
SubspeciesLaudakia nupta fusca (BLANFORD 1876)
Laudakia nupta nupta (DE FILIPI 1843) 
Common Namesnupta: Large-scaled (Rock) Agama
fusca: Yellow-headed Rock Agama 
SynonymAgama nupta DE FILIPI 1843: 407
Stellio nuptus — BLANFORD 1876: 317
Stellio carinatus DUMÉRIL 1851 (fide SMITH 1935)
Agama nupta — BOULENGER 1885: 365
Agama flavicauda WERNER 1897: 264 (fide WERMUTH 1967)
Agama flavicauda — LOVERIDGE 1920: 141
Agama nupta — SMITH 1935: 219
Agama caucasica SCHMIDT 1939
Agama nupta nupta — MINTON 1966: 475
Agama nupta nupta — WERMUTH 1967: 20
Laudakia nupta — FRYNTA et al. 1997
Laudakia nupta — BARTS & WILMS 2003
Laudakia nupta nupta — BAIG et al. 2012: 245

Laudakia nupta fusca (BLANFORD 1876)
Stellio nuptus var. fuscus BLANFORD 1876: 319
Agama nupta var. fusca — BOULENGER 1885: 366
Agama nupta fusca — MINTON 1966: 92
Agama nupta fusca — WERMUTH 1967: 20
Laudakia fusca — CHEATSAZAN et al. 2008
Laudakia nupta fusca — BAIG et al. 2012 
Distributionnupta: E Iraq, Iran, Afghanistan, Pakistan; Type locality: Persepolis [Fars province, Iran].

fusca: S Pakistan, S Iran; Type locality: Kalagán (elevation 3500 feet) and near Jálk (3000 feet), Baluchistan. BAIG et al. 2012 gave “Iran, Sistan-Baluchistan Province near Jalk (3000 f)” as type locality. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesSyntypes: MSNTO R322 (Museo Civico di Storia Naturale, Torino); ZSI 6798, ZMB 6078, "Sciraz, Persien", leg. G. Doria.
Lectotype: BMNH 74.11.32.11 (designated by RASTEGAR-POUYANI & NILSON 2002)
Syntypes: ZMB 13723, ZMB 54608-09 (was ZMB 13723 A+B), without locality, leg. Schweinfurth [flavicauda] 
CommentSynonymy mostly after WERMUTH 1967. BAIG (1992) considers L. fusca as a synonym of L. nupta, Agama nupta fusca has been elevated to full species status by CHEATSAZAN et al. 2008, but was relegated to subspecies status by BAIG et al. 2012. SMITH (1935) did not find any significant difference between L. n. nupta, L. n. fusca and Stellio carinatus and thus placed all three together under L. nupta.

Diagnosis (fusca). Size large; head heavy; nostril on canthus rostralis; body moderately depressed; absence of a nuchal fold across the neck; vertebral enlarged scales strongly mucronate and cover most of dorsal part of body; no patch of enlarged mucronate scales on flanks; tail long and has segments of three whorls; callous glands present mostly in males [BAIG et al. 2012].

Diagnosis (nupta). Size large; head heavy; nostril on can- thus rostralis; body moderately depressed; yellowish or reddish brown; vertebral enlarged scales strongly keeled and cover most of dorsal part of body; no patch of enlarged mucronate scales on flanks; tail long and has segments of three whorls; callous glands present mostly in males [BAIG et al. 2012]. 
References
  • Ananjeva, N.B. & Tuniev 1994. Some aspects of historical biogeography of Asian rock agamids. Russ. J. Herpetol. 1 (1): 43 - get paper here
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144]
  • Anderson,S.C. 1963. Amphibians and Reptiles from Iran. Proc. Cal. Acad. Sci. Ser. 4, 31(16): 417-498 - get paper here
  • Baig, Khalid Javed; PhiliPP Wagner, Natalia B. AnanJeva & Wolfgang Böhme 2012. A morphology-based taxonomic revision of Laudakia Gray, 1845 (Squamata: Agamidae). Vertebrate Zoology 62 (2): 213-260 - get paper here
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Blanford,W.T. 1876. Eastern Persia, an account of the journeys of the Persian Boundary Comission, 1870-71-72. Vol. II. The zoology and geology. London, 516 pp.
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  • Boulenger, George A. 1890. The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia. Taylor & Francis, London, xviii, 541 pp. - get paper here
  • Cheatsazan, Hamed; Vahideh Rabani, Atabak Mahjoorazad and Haji Gholi Kami 2008. Taxonomic status of the Yellow-Headed Agama, Laudakia nupta fusca (Blanford, 1876) (Sauria: Agamidae). Zoology in the Middle East 44: 41-50 - get paper here
  • Clark, R. J., Clark, E. D., Anderson, S. C. and Leviton, A. E. 1969. Report on a collection of amphibians and reptiles from Afghanistan. Proc. Cal. Acad. Sci. (ser. 4) 36: 279-316. - get paper here
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  • Frynta,D. et al. 1997. Results of the Czech biological expedition to Iran. Part 1. Notes on the distribution of amphibians and reptiles. Acta Soc. Zool. Bohem. 61: 3-17
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  • Loveridge, A. 1923. Notes on East African lizards collected 1920-1923 with the description of two new races of Agama lionotus Blgr. Proc. Zool. Soc. London 1923: 935-969 - get paper here
  • Loveridge,A. 1920. Notes on East African lizards collected 1915-1919, with description of a new genus and species of skink and new subspecies of gecko. Proc. Zool. Soc. London 1920: 131-167 - get paper here
  • Macey, J. R., J. A. Schulte II, A. Larson, N. B. Ananjeva, Y. Wang, R. Pethiyagoda, N. Rastegar-Pouy 2000. Evaluating trans-Tethys migration: an example using acrodont lizard phylogenetics. Systematic Biology 49 (2): 233-256 - get paper here
  • Minton, S. A. Jr. 1966. A contribution to the herpetology of West Pakistan. Bull. Amer. Mus. Nat. Hist., 134: 29-184.
  • Nikolsky, A. M. 1897. Les reptiles, amphibiens et poissons recueillis (part.) Mr. N. Zaroudny dans la Perse orientale [in Russian]. Annuaire Musée Zoologique de l’Académie Impériale des Sciences de St.-Pétersbourg 2: 306-348
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT A 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics4 (1): 7-30
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  • Werner, F. 1917. Reptilien aus Persien (Provinz Fars). Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 67:191—220.
 
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