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Laudakia tuberculata (GRAY, 1827)

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards) 
Subspecies 
Common NamesTuberculated Agama, Kashmir Rock Agama 
SynonymAgama tuberculata GRAY 1827: 218
Agama tuberculata — DUMÉRIL & BIBRON 1837: 488
Stellio indicus BLYTH 1853: 646
Stellio tuberculatus — GÜNTHER 1864
Agama tuberculata — BOULENGER 1885: 361
Agama tuberculata — SMITH 1935: 214
Agama tuberculata — MINTON 1966
Agama tuberculata — SURA 1989
Laudakia tuberculata — DAS 1996: 44
Laudakia tuberculata — MACEY et al. 2000
Laudakia tuberculata — BAIG et al. 2012: 251
Laudakia tuberculata — BAIG et al. 2012: 25 
DistributionN Pakistan
India (W Himalaya, Kashmir, Punjab, Jammu, Uttar Pradesh, Himachal Pradesh),
Nepal (Katmandu, Chitlong), E Afghanistan
China (Xizang = Tibet)

According to BAIG et al. 2012 in “E Afghanistan, NW Pakistan, Kashmir, Uttar Pradesh, India, SW Nepal.”

Type locality: India; restricted to Bengal by SMITH 1935.

indicus: Type locality: “Mirzapore [= Wazirabad, upper Hindustan].”  
Reproductionoviparous 
TypesSynpes: BMNH 1946.8.28.17/60.3.19.1377 
DiagnosisDiagnosis (genus). Head and body depressed; head scales smooth or rugose; no gular pouch; usually no patch of enlarged or spinose scales on flanks (but present in some species); gular scales smooth; ventral scales smooth; vertebral scales enlarged, sometimes with median rows of small scales; groups of spinose scales on neck and sides of head; tail segment of two to four whorls, usually three to four [BAIG et al. 2012].
 
CommentSynonymy: Stellio dayanus STOLIZKA 1871 has been removed from the synonymy of L. tuberculata by BAIG & BÖHME (1997). Synonymy mostly after WERMUTH 1967.

Type species: Agama tuberculata GRAY 1827 is the type species of the genus Laudakia GRAY 1845: 254.

Type species: Barycephalus sykesii Günther = Agama tuberculata HARDWICKE & GRAY, 1827) is the type species of the genus Barycephalus Günther (non Barycephalus Brauns, 1895 [= Hymenoptera]), now a synonym of Laudakia. 
Etymology 
References
  • Ananjeva, N.B. & Tuniev 1994. Some aspects of historical biogeography of Asian rock agamids. Russ. J. Herpetol. 1 (1): 43 - get paper here
  • ANANJEVA, Natalia B.; Xianguang GUO and Yuezhao WANG 2011. Taxonomic Diversity of Agamid Lizards (Reptilia, Sauria, Acrodonta, Agamidae) from China: A Comparative Analysis. Asian Herpetological Research 2 (3): 117-128 - get paper here
  • Bahuguna, Archana. 2008. Altitudinal variations in morphological characters of Laudakia tuberculata Hardwicke, and Gray, 1827 from western Himalayas (Uttarakhand), India. Russ. J. Herpetol. 15 (3):207-211 - get paper here
  • Baig, Khalid Javed; Philipp Wagner, Natalia B. Ananjeva & Wolfgang Böhme 2012. A morphology-based taxonomic revision of Laudakia Gray, 1845 (Squamata: Agamidae). Vertebrate Zoology 62 (2): 213-260 - get paper here
  • Baig,K.J. & Böhme,W. 1995. Partition of the Stellio-group of Agama. 8th Ord. Gen. Meet. Soc. Europ. Herpet.: 36
  • Blyth,E. 1854. Notices and descriptions of various reptiles, new or little-known. Part I. J. Asiat. Soc. Bengal 22 [1853]: 639-655 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger, G.A., Annandale, N., Wall, F. & Regan, C.T. 1907. Reports on a collection of batrachia, reptiles and fish from Nepal and the western Himalayas. Lacertilia. Records of the Indian Museum 1:149—155 - get paper here
  • Boulenger, George A. 1890. The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia. Taylor & Francis, London, xviii, 541 pp. - get paper here
  • Dodsworth, P.T.L. 1913. On the Habits of the Rock Lizard (Agama tuberculata). J. Bombay nat. Hist. Soc. 22: 404 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Fischer, C.E.C. 1907. Aberration in Scales of regrown tail of Agama tuberculata, Grey. J. Bombay Nat. Hist. Soc. 18: 208 - get paper here
  • Gray. J. E. 1834. Illustrations of Indian Zoology, chiefly selected from the collection of Major - General Hardwicke. Vol. 2. London (1833-1834): 263 pp., 95 plates - get paper here
  • Günther, A. 1864. The Reptiles of British India. London (Taylor & Francis), xxvii + 452 pp. - get paper here
  • Günther,A. 1860. Contributions to a knowledge of the reptiles of the Himalaya mountains. - I. Descriptions of the new species. II. List of Himalayan reptiles, with remarks on their horizontal distribution. Proc. Zool. Soc. London 1860: 148-175 - get paper here
  • Hardwicke, Thomas & Gray, J.E. 1827. A synopsis of the species of saurian reptiles, collected in India by Major-General Hardwicke. Zoological Journal, London 3: 213-229 - get paper here
  • Kästle , W., Rai, K. & Schleich, H.H. 2013. FIELD GUIDE to Amphibians and Reptiles of Nepal. ARCO-Nepal e.V., 625 pp. - get paper here
  • Khan, B.; Ahmed, W.; Ablimit, A.; Fakhri, S.; Ali, H. 2012. Range extension of four highland agamid lizards in Shimshal Pamir, Pakistan. J. Arid Land 4 (1): 77-84
  • Kumar, Amit, Shruti Sengupta, Upma Manral and Abhijit Das. 2016. Laudakia tuberculata (Kashmir Rock Agama) Morphological anomaly. Herpetological Review 47 (2): 300
  • Macey, J. R., J. A. Schulte II, A. Larson, N. B. Ananjeva, Y. Wang, R. Pethiyagoda, N. Rastegar-Pouyani, T. J. Papenfuss 2000. Evaluating trans-Tethys migration: an example using acrodont lizard phylogenetics. Systematic Biology 49 (2): 233-256 - get paper here
  • Minton, S. A. Jr. 1966. A contribution to the herpetology of West Pakistan. Bull. Amer. Mus. Nat. Hist., 134: 29-184.
  • Murthy, T.S.N. 2010. The reptile fauna of India. B.R. Publishing, New Delhi, 332 pp.
  • Nanhoe, L.M.R., & Ouboter, P.E. 1987. The distribution of reptiles and amphibians in the Annapurna-Dhaulagiri region (Nepal). Zoologische Verhandelingen (240): 1-105 - get paper here
  • Saikia, U.; Sharma, D.K. & Sharma, R.M. 2007. Checklist of the Reptilian fauna of Himachal Pradesh, India. Reptile Rap (8): 6-9 - get paper here
  • Schmidt,K.P. 1926. Amphibians and Reptiles of the James Simpson-Roosevelt Asiatic Expedition. Field Mus. nat. Hist., Zool. Ser. 12: 167-173 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Singh, Vikram; H. S. Banyal 2014. First Record of Kashmir Rock Agama (Laudakia tuberculata Hardwicke et Gray, 1827) from Kalatop-Khajjiar Wildlife Sanctuary, Chamba (Himachal Pradesh), India. Russ. J. Herpetol. 21 (3): 234-236 - get paper here
  • Smith,M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Sura, P. 1989. Über eine kleine Sammlung von Fröschen und Echsen aus dem zentralen Teil Nepals. Herpetofauna 11 (58): 28-31 - get paper here
  • Swan, L.W., & Leviton, A.E. 1962. The herpetology of Nepal: a history, check list, and zoogeographical analysis of the herpetofauna. Proc. Cal. Acad. Sci. 32 (6) (4.s.): 103-147. - get paper here
  • Venugopal, P.D. 2010. An updated and annotated list of Indian lizards (Reptilia: Sauria) based on a review of distribution records and checklists of Indian reptiles. Journal of Threatened Taxa 2 (3): 725-738. - get paper here
  • Waltner, R.C. 1975. Geographical and altitudinal distribution of amphibians and reptiles in the Himalayas. Cheetal (Dehra Dun, India) 16(1): 17-25; 16(2): 28-36; 16(3): 14-19; 16(4): 12-17.
 
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