Leiocephalus macropus (COPE, 1863)

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Higher Taxa	Leiocephalidae, Iguania, Sauria, Squamata (lizards)
Subspecies	Leiocephalus macropus macropus COPE 1863 Leiocephalus macropus aegialus SCHWARTZ & GARRIDO 1967 Leiocephalus macropus asbolomus SCHWARTZ & GARRIDO 1967 Leiocephalus macropus felinoi GARRIDO 1979 Leiocephalus macropus hoplites ZUG 1959 Leiocephalus macropus hyacinthurus ZUG 1959 Leiocephalus macropus immaculatus HARDY 1958 Leiocephalus macropus koopmani ZUG 1959 Leiocephalus macropus lenticulatus GARRIDO 1973 Leiocephalus macropus phylax SCHWARTZ & GARRIDO 1967 Leiocephalus macropus torrei GARRIDO 1979
Common Names	E: Cuban Side-blotched Curlytail, Monte Verde Curlytail Lizard
Synonym	Liocephalus (sic) macropus COPE 1863:184 Liocephalus macropus — BOULENGER 1885: 163 Leiocephalus macropus — STEJNEGER 1917 Leiocephalus macropus — SCHWARTZ & HENDERSON 1991: 435 Leiocephalus macropus — RODRÍGUEZ SCHETTINO 1999: 135 Leiocephalus macropus aegialus SCHWARTZ & GARRIDO 1967 Leiocephalus macropus aegialus — PREGILL 1992: 47 Leiocephalus macropus aegialus — HEDGES et al. 2019 Leiocephalus macropus asbolomus SCHWARTZ & GARRIDO 1967 Leiocephalus macropus asbolomus — PREGILL 1992: 47 Leiocephalus macropus asbolomus — HEDGES et al. 2019 Leiocephalus macropus felinoi GARRIDO 1979 Leiocephalus macropus felinoi — PREGILL 1992: 47 Leiocephalus macropus felinoi — HEDGES et al. 2019 Leiocephalus macropus hoplites ZUG 1959 Leiocephalus macropus hoplites — PREGILL 1992: 47 Leiocephalus macropus hoplites — HEDGES et al. 2019 Leiocephalus macropus hyacinthurus ZUG 1959 Leiocephalus macropus hyacinthurus — PREGILL 1992: 47 Leiocephalus macropus hyacinthurus — HEDGES et al. 2019 Leiocephalus macropus immaculatus HARDY 1958 Leiocephalus macropus immaculatus — PREGILL 1992: 47 Leiocephalus macropus inmaculatus [sic] — RODRÍGUEZ SCHETTINO 1999: 137 Leiocephalus macropus immaculatus — HEDGES et al. 2019 Leiocephalus macropus koopmani ZUG 1959 Leiocephalus macropus koopmani — PREGILL 1992: 47 Leiocephalus macropus koopmani — HEDGES et al. 2019 Leiocephalus macropus lenticulatus GARRIDO 1973: 10 Leiocephalus macropus lenticulatus — PREGILL 1992: 47 Leiocephalus macropus lenticulatus — HEDGES et al. 2019 Leiocephalus macropus phylax SCHWARTZ & GARRIDO 1967 Leiocephalus macropus phylax — PREGILL 1992: 47 Leiocephalus macropus phylax — HEDGES et al. 2019 Leiocephalus macropus torrei GARRIDO 1979 Leiocephalus macropus torrei — PREGILL 1992: 47 Leiocephalus macropus torrei — HEDGES et al. 2019
Distribution	Cuba (from Pinar del Rio east, but primarily E Cuba) macropus: SE coast from Bahia de Santiago east at least to Sierra del Purial north of Cajobabo; presumably inland to type-locaiity in Sierra del Guaso; Isla de la Juventud? Type locality: Eastern Cuba; restricted by Stejneger, 1917, to Monte Verde, Guantánamo Province, Cuba. aegialus: northern coastal Camagüey Province. asbolomus: northeastern coast of Guantânamo Province, west to Holguin Province. felinoi: north coastal Matanzas Province. hoplites: northern Ciego de Âvila Province. hyacinthurus: Sierra de Escambray. immaculatus: southeastern coast from Rio Magdalena east to Bahia de Santiago; Cayo Damas. koopmani: Peninsula de Guanahacabibes. lenticulatus: coastal hills west of Gibara, Holguin Province. Type locality: “arenal Los Cocos, a seis km de Gibara” phylax: southwestern Granma Province, from Verre6n east to Rio Puercos and Punta Hicacos. torrei: along Rio Paredones, Matanzas Province. INCERTAE SEDIS: Material from the following localities has not been assigned to subspecies: Niquero, Cayo Saetia, Pinares de Mayari, Sierra de Nipe, Cupeyai, all in eastern Cuba; Miranda, Santiago de Cuba Province; Rangel, Pinar del Rio Province; Punta Hicacos, Matanzas Province.
Reproduction	oviparous
Types	Lectotype: USNM 25819, selected by Hardy, 1958; Syntypes: MCZ 10930; USNM 12254; USNM 25819-23, USNM 25825-26. Holotype: AMNH 83255 [aegialus] Holotype: CZACC = IZ 568 [asbolomus] Holotype: CZACC = IZ 4751 [felinoi] Holotype: AMNH 78020 [hoplites] Holotype: AMNH 78015 [hyacinthurus] Holotype: USNM 138412 [immaculatus] Holotype: MCZ 55541 [koopmani] Holotype: CZACC = IZ 2782 [lenticulatus] Holotype: CZACC = IZ 556 [phylax] Holotype: CZACC = IZ 3725 [torrei]
Diagnosis
Comment	See Schwartz and Garrido, 1967, for comments on restricted type-locality, status of syntypes, and selection of lectotype. Distribution: see map of subspecies in RODRÍGUEZ SCHETTINO 1999: 136.
References	Alfonso, Yasel U.; Pierre Charruau, Gabriel Fajardo and Alberto R. Estrada. 2012. Interspecific communal oviposition and reproduction of three lizard species in Southeastern Cuba. Herpetology Notes 5: 73-77. Bakkegard, Kristin A.; David A. Johnson, and Daniel G. Mulcahy 2020. A New Locality, Naval Station Guantanamo Bay, for the Rare Lizard Leiocephalus onaneyi (Guantánamo Striped Curlytail) and Notes on its Natural History. CARIBBEAN NATURALIST (79): 1–22 Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here Cope, E.D. 1863. Contributions to Neotropical saurology. Proc. Acad. Nat. Sci. Philadelphia 14 [1862]: 176-188. - get paper here Garrido, O. H. 1973. Nueva especie de Leiocephalus (Lacertilia, Iguanidae) para Cuba. Poeyana 116:1-19. Garrido, O.H. 1973. Nuevas subespecies de reptiles para Cuba. Torreia, n.s. 30:1-31. Garrido, O.H. 1979. Nuevas subespecies de Leiocephalus macropus Cope (Lacertilia: Iguanidae) para Cuba. Poeyana (188):1-16. Hardy, J.D., Jr. 1958. A geographic variant gradient in the Cuban lizard Leiocephalus macropus Cope. Herpetologica 13 (3): 275-76 - get paper here Hedges SB, Powell R, Henderson RW, Hanson S, and Murphy JC 2019. Definition of the Caribbean Islands biogeographic region, with checklist and recommendations for standardized common names of amphibians and reptiles. Caribbean Herpetology 67: 1–53 Pregill,G.K. 1992. Systematics of the West Indian Lizard Genus Leiocephalus (Squamata: Iguania: Tropiduridae). Misc. Publ. Univ. Kans. Mus. Nat. Hist. 84: 1-69 - get paper here Rodríguez Schettino, Lourdes (ed.) 1999. The iguanid lizards of Cuba. University Press of Florida, 428 pp. Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp. Schwartz, A. and Garrido, O.H. 1967. A review of the Cuban lizard Leiocephalus macropus Cope. Reading Public Museum and Art Gallery Sci. Publ. (14): 1-41. Zug, G.R. 1959. Three new subspecies of the lizard Leiocephalus macropus Cope from Cuba. Proc. Biol. Soc. Washington72:139-50. - get paper here
External links	IUCN Red List National Center for Biotechnology Information http://fm2.fieldmuseum.org/plantguides/guide_pdfs/197%20Cuba%20Baconao… http://2.bp.blogspot.com/-1MwwViYJSIM/TsK0T_vonQI/AAAAAAAAUqI/Fl_DLfV_QpA… Google images

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