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Letheobia pembana BROADLEY & WALLACH, 2007

IUCN Red List - Letheobia pembana - Data Deficient, DD
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Higher TaxaTyphlopidae (Afrotyphlopinae), Typhlopoidea, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Pemba gracile blind-snake 
SynonymLetheobia pembana BROADLEY & WALLACH 2007: 45
Typhlops pallidus — (not COPE) BOETTGER 1913: 351
Typhlops pallidus — BOULENGER 1915: 616 (part)
Typhlops pallidus — MOREAU & PAKENHAM 1941: 108 (part);
Typhlops pallidus — LOVERIDGE 1957: 244 (part)
Rhinotyphlops pallidus – (not COPE) PAKENHAM 1983: 21 (part)
Rhinotyphlops pallidus – SPAWLS et al. 2002: 295 (part)
Letheobia pembana — HEDGES et al. 2014
Letheobia pembana — SPAWLS et al. 2018: 359
Letheobia pembana — WALLACH et al. 2014: 381 
DistributionTanzania: Pemba Island

Type locality: Pemba Island, Tanzania (05°15’S, 39°50’E, elevation <100 m).  
Reproductionoviparous 
TypesHolotype: SMF 16688, a male, collected by A. Voeltzkow in 1905. 
DiagnosisDiagnosis: A member of the Letheobia pallida complex, but distinguished by its more numerous scale rows (26-24-24 vs 24-22-22) and lower number of middorsals (353 vs 376-487).
Description: Snout rounded, prominent. Rostral very broad, truncated posteriorly; frontal crescentic; supraocular transverse, its lateral apex between nasal and ocular, the latter separated from the lip by a large subocular; eye not visible; nasal suture arising from second labial; SIP X (N1, P, O, O); scale rows 26-24-24; MD 353; L/D ratio 53. Length 158 mm. Colourless. (BROADLEY & WALLACH 2007: 45) 
CommentHabitat: Coastal mosaic, probably cultivation steppe. 
EtymologyNamed for Pemba Island, the type locality. 
References
  • Boettger, O. 1913. Reptilien und Amphibien von Madagascar, den Inseln und dem Festland Ostafrikas. Pp. 269-375. In: Voeltzkow, A. Reise in Ostafrika in den Jahren 1903-1905. Wissenschaftliche Ergebnisse. Vol. 3. Systematische Arbeiten. Schweizerbart’ sche Verlagsbuchhandlung, Nägele und Sproesser, Stuttgart - get paper here
  • Boulenger, G.A. 1915. A list of the snakes of East Africa, north of the Zambesi and south of the Soudan and Somaliland, and of Nyasaland. Proc. Zool. Soc. London,1915, 611–640 - get paper here
  • Broadley, Donald G. & Wallach, V. 2007. A review of East and Central African species of Letheobia Cope, revived from the synonymy of Rhinotyphlops Fitzinger, with descriptions of five new species (Serpentes: Typhlopidae). Zootaxa 1515: 31–68 - get paper here
  • Hawlitschek, O.; F. Glaw & D. Rödder 2012. Pemba – Herpetologische Fundgrube im Indischen Ozean. Reptilia (Münster) 17 (97): 97-109 - get paper here
  • Hedges, S.B., Marion, A.B., Lipp, K.M., Marin, J. & Vidal, N. 2014. A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology 49: 1–61 - get paper here
  • Loveridge, A. 1957. Check list of the reptiles and amphibians of east Africa (Uganda, Kenya, Tanganyika, Zanzibar). Bull. Mus. Comp. Zool. Harvard 117 (2): 153-362 - get paper here
  • Moreau, R.E., & Pakenham, R.H.W. 1940. The land vertebrates of Pemba, Zanzibar, and Mafia: a zoogeographical study. Proc. Zool. Soc. London (Ser. A) 110: 97-128.
  • Pakenham, R.H.W. 1983. The reptiles and amphibians of Zanzibar and Pemba Islands (with a note on the freshwater fishes). J. of the East Afr. Natur. Hist. Soc. and Natl. Mus. no. 177 40 pp.
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Wallach, V. & Gemel, R. 2018. Typhlops weidholzi n. inedit., a new species of Letheobia from the republic of Cameroon, and a synopsis of the genus (Squamata: Serpentes: Scolecophidia: Typhlopidae). Herpetozoa 31 (1/2): 27 - 46 - get paper here
 
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