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Liolaemus schmidti (MARX, 1960)

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Higher TaxaLiolaemidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Schmidt's Tree Iguana

E: Panther Tree Iguana [islugensis]
S: Lagartija pantera [islugensis] 
SynonymCtenoblepharys schmidti MARX 1960
Liolaemus schmidti — CEI 1979
Liolaemus islugensis ORTIZ & MARQUET 1987
Liolaemus schmidti — DIRKSEN & DE LA RIVA 1999
Liolaemus islugensis — LOBO et al. 2010
Liolaemus islugensis — ABDALA et al. 2013
Liolaemus schmidti — ABDALA et al. 2013
Liolaemus islugensis — DEMANGEL 2016
Liolaemus schmidti — LANGSTROTH 2021 
DistributionChile (Andes of Antofagasta between Chile and Bolivia); Bolivia (Bolivia, Potosí)

Type locality: Forty miles east of San Pedro de Atacama, Antofagasta, Chile.

islugensis (invalid): Chile (Iquique), Bolivia (Potosí, Tarija); Type locality: Colchane 3,850 m elevation, Provincia de Iquique, Chile (19° 42’ S, 68° 53’ W).  
Reproductionviviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: FMNH 5759.
Holotype: MZUC 10931, male (Museo de Zoología de la Universidad de Concepción, Chile) [islugensis] 
Diagnosis 
CommentSynonymy: PINCHEIRA-DONOSO & NÚÑEZ 2005 and PINCHEIRA-DONOSO et al. 2008 considered L. schmidti as a synonym of L. andinus, a decision that is rejected by LOBO et al. 2010. More recently, PINCHEIRA-DONOSO agreed that schmidti is a valid species (pers. comm., 23 May 2017). PINCHEIRA-DONOSO & NUNEZ (2003) synonymized L. islugensis with L. pantherinus. Langstroth 2021 synonymized islugensis with L. schmidti.

Subspecies: Liolaemus islugensis erguetae LAURENT 1995 has been elevated to species status.

Distribution: not listed for Chile by Demangel 2016, apparently because he considered it as a synonym of something else. See map in Langstroth 2021: 124 (Fig. 6).

NCBI taxonID: 2316808 [islugensis]

RDB species ID: 20949 [islugensis]

Etymology: L. islugensis was named after Volcán Isluga, where this species is found. 
EtymologyNamed after Karl Patterson Schmidt (1890-1957), American herpetologist who died from a Boomslang bite (Dispholidus typus; see Davis 1957 and RESETAR & VORIS 1997 for biographical notes). 
References
  • Abdala CS, Aguilar-Kirigin AJ, Semhan RV, Bulacios Arroyo AL, Valdes J, Paz MM, et al. 2019. Description and phylogeny of a new species of Liolaemus (Iguania: Liolaemidae) endemic to the south of the Plurinational State of Bolivia. PLoS ONE 14(12): e0225815 - get paper here
  • Abdala, C.S.; S. Quinteros & R.E. Espinoza 2008. Two new species of Liolaemus (iguania: Liolaemidae) from the Puna of Northwestern Argentina. Herpetologica 64 (4): 458–471 - get paper here
  • Abdala, Cristian Simón; Juarez Heredia, Viviana Isabel 2013. Taxonomía y filogenia de un grupo de lagartos amenzados: el grupo de Liolaemus anomalus (Iguania:Liolaemidae). Cuadernos de Herpetología 27 (2): - get paper here
  • Abdala, Cristian Simón; Marcos Maximiliano Paz & Romina Valeria Semhan 2013. Nuevo Liolaemus (Iguania: Liolaemidae) con novedoso carácter morfológico, de la frontera entre Argentina y Chile. Rev. Biol. Trop. 61 (4): 1563-1584 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Cei ,J.M. 1979. A reassessment of the genus Ctenoblepharis (Reptilia, Sauria, Iguanidae) with description of a new subspecies of Liolaemus multimaculatus from western Argentina. Journal of Herpetology 13 (3): 297-302 - get paper here
  • Davis, D.D. 1959. Karl Patterson Schmidt, 1890-1957. Copeia 1959 (3): 189-192 - get paper here
  • Demangel, Diego 2016. Reptiles en Chile. Fauna Nativa Ediciones, Santiago, 619 pp - get paper here
  • Dirksen, L. & De la Riva, I. 1999. The lizards and amphisbaenians of Bolivia (Reptilia, Squamata): checklist, localities, and bibliography. Graellsia 55: 199-215 - get paper here
  • Langstroth, Roberto Plotkin 2021. The identities of Liolaemus signifer (Duméril & Bibron 1837), L. pantherinus Pellegrin 1909, L. schmidti (Marx 1960), and L. erroneus (Núñez & Yáñez 1984 “1983-1984”) (Squamata: Liolaemidae). Cuadernos de Herpetología 35 (S1): 111-168
  • Lobo, F.; Espinoza, R.E. and S. Quinteros 2010. A critical review and systematic discussion of recent classification proposals for liolaemid lizards. Zootaxa 2549: 1–30 - get paper here
  • Marx,H. 1960. A new iguanid lizard of the genus Ctenoblepharis. Fieldiana Zool. 39: 407-409 - get paper here
  • Núñez, H. and A. Veloso 2001. Distribución geográfica de las especies de lagartos de la región de Antofagasta, Chile. Bol. Mus. Nac.Hist. Nat. 50: 109-120
  • Ortiz J C; Marquet P 1987. Una nueva especie de lagarto altoandino: Liolaemus islugensis (Reptilia-Iguanidae). GAYANA ZOOLOGIA 51 (1-4): 59-63 - get paper here
  • Yáñez, J. and H. Núñez. 1982. Segundo registro de Liolaemus schmidti (Marx, 1960). (Lacertilia: Iguanidae). Noticiario Mensual del Museo Nacional de Historia Natural (Chile) 26(303-304): 12
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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