Luperosaurus cumingii (GRAY, 1845)

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Higher Taxa	Gekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies
Common Names	E: Philippine Wolf Gecko
Synonym	Lyperosaurus cumingii GRAY 1845: 145 Luperosaurus cumingii GRAY 1845: 163 Luperosaurus cumingii — BOULENGER 1885: 181 Luperosaurus cumingii — TAYLOR 1922: 86 Luperosaurus cumingii — WERMUTH 1965: 101 Luperosaurus cumingii — KLUGE 1993 Luperosaurus cumingi — BROWN et al. 2000 Luperosaurus cf. cumingii — GAULKE et al. 2003 Luperosaurus cumingii — OLIVER et al. 2018
Distribution	Philippines (S Luzon) Type locality: Philippines.
Reproduction	oviparous
Types	syntypes: BMNH 1946.8.22.41 and 1946.8.22.42;
Diagnosis	Diagnosis (genus): The genus Luperosaurus is characterized by a unique combination of morphological features, including moderately to strongly developed cutaneous folds on the posterior and, in some species, anterior border of the limbs, broadly dilated digits, a lack of enlarged postmental scales, small scales encircling the tail (differentiated subcaudals absent), and well-developed interdigital webbing (Gaulke et al. 2007). According to DAS et al. (2008), the genus is characterized by the following characters: chin shields undifferentiated; digits dilated, half-webbed; first digit clawless; cutaneous lobes present on posterior margins of limbs; and subcaudals undifferentiated (R. M. BROWN et al., 2000; 2007; RUSSELL 1979).
Comment	Type species: Luperosaurus cumingii GRAY 1845 is the type species of the genus Luperosaurus GRAY 1845. Das (2005), as First Reviser, selected the nomen Luperosaurus Gray, 1845 as valid for the taxon, instead of Lyperosaurus. Phylogeny: Luperosaurus is nested within Lepidodactylus s.l. (fide Oliver et al. 2018 who did not make any formal changes to the nomenclature). Distribution: the range of L. cumingii was restricted to S Luzon by BROWN et al. who described the other populations from Panay and Negros as a new species, L.
Etymology	Named after Hugh Cuming (1791-1865), an English naturalist and conchologist. See Beolens et al. 2011: 60 for more details.
References	Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here Brown, R. M. and A. C. Diesmos 2000. The lizard genus Luperosaurus: taxonomy, history, and conservation prospects for some of the world's rarest lizards. Sylvatrop: Technical Journal of Philippine Ecosystems and Natural Resources 10:107-124. Brown, R.M., Diesmos, A.C., Duya, M.V., Garcia, H.J.D., & Rico, E.L. 2010. New forest gecko (Squamata; Gekkonidae; Genus Luperosaurus) from Mt. Mantalingajan, southern Palawan Island, Philippines. Journal of Herpetology 44 (1): 37–48 - get paper here Brown, R.M.; Diesmos, A. C. & Duya, M.V. 2007. A new Luperosaurus (Squamata: Gekkonidae) from the Sierra Madre of Luzon Island, Philippines. Raffles Bulletin of Zoology 55 (1): 167-174 - get paper here Brown, Rafe M., Jatna Supriatna and Hidetoshi Ota. 2000. Discovery of a new species of Luperosaurus (Squamata: Gekkonidae) from Sulawesi, with a phylogenetic analysis of the genus, and comments on the status of Luperosaurus serraticaudus. Copeia 2000 (1): 191-209 - get paper here Brown, Rafe M.; Arvin C. Diesmos, and Carl H. Oliveros 2011. New Flap-Legged Forest Gecko (Genus Luperosaurus) from the Northern Philippines. Journal of Herpetology 45 (2): 202-210. - get paper here Das, I. 2005. Nomenclatural Notes on the Generic Nomen Luperosaurus Gray, 1845 (Squamata: Gekkonidae). Herpetological Review 36 (2): 117–118 - get paper here Das, Indraneil; Lakim, Maklarin; Kandaung, Pius. 2008. New species of Luperosaurus (Squamata: Gekkonidae) from the Crocker Range Park, Sabah, Malaysia (Borneo). Zootaxa 1719: 53-60 - get paper here Gaulke, M., H. Rösler & R. M. Brown 2007. A new species of Luperosaurus (Squamata: Gekkonidae) from Panay Island, Philippines, with comments on the taxonomic status of Luperosaurus cumingii (Grey, 1845). Copeia 2007 (2):413-425 - get paper here Gaulke, M.; Demegillo, A.; Reiter, J. & Tacud, B. 2003. Additions to the herpetofauna of Panay Island, Philippines. Salamandra 39 (2): 111-122 - get paper here Gojo-Cruz, Paul Henric P. and Leticia E. Afuang 2018. The Zoogeographic Significance of Caraballo Mountain Range, Luzon Island, Philippines With Focus on the Biogeography of Luzon’s Herpetofauna. Philippine Journal of Science 147 (3): 393-409 - get paper here Gray, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp. - get paper here Oliver, Paul M.; Rafe M. Brown, Fred Kraus, Eric Rittmeyer, Scott L. Travers, Cameron D. Siler 2018. Lizards of the lost arcs: mid-Cenozoic diversification, persistence and ecological marginalization in the West Pacific. Proc. R. Soc. B 285 20171760; DOI: 10.1098/rspb.2017.1760 - get paper here Rafe M. Brown, Cameron D. Siler, Indraneil Das, Yong Min 2012. Testing the phylogenetic affinities of Southeast Asia’s rarest geckos: Flap-legged geckos (Luperosaurus), Flying geckos (Ptychozoon) and their relationship to the pan-Asian genus Gekko. Molecular Phylogenetics and Evolution 63: 915-921 - get paper here Russell A P 1979. A new species of Luperosaurus (Gekkonidae) with comments on the genus. Herpetologica 35 (3): 282-288 - get paper here Taylor, E.H. 1922. The lizards of the Philippine Islands. Department of Agriculture and Natural Resources, Bureau of Science, Government of the Philippine Islands, Manila, Publication no. 17: 269 pp. - get paper here
External links	IUCN Red List National Center for Biotechnology Information Google images

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