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Lycodon albofuscus (DUMÉRIL, BIBRON & DUMÉRIL, 1854)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Dark Wolf Snake
G: Dunkle Wolfszahnnatter 
SynonymSphecodes albo-fuscus DUMÉRIL, BIBRON & DUMÉRIL 1854: 394
Sphecodes albo-fuscus DUMÉRIL 1853 (nom. nud., generic name not latinized)
Sphecodes albofuscus — BLEEKER 1858
Ophites albofuscus — GÜNTHER 1858
Ophites albofuscus — BOULENGER 1885: 389
Lepturophis borneensis BOULENGER 1900: 183
Lycodon albofuscus — BOULENGER 1893: 357
Lepturophis borneensis — GRANDISON 1972: 87
Lepturophis borneensis — GRANDISON 1978
Lycodon albofuscus — MANTHEY 1983
Lepturophis borneensis — MURPHY et al. 1994
Lepturophis albofuscus — MANTHEY & GROSSMANN 1997: 359
Lepturophis borneensis — COX et al. 1998: 64
Lepturophis albofuscus — CHAN-ARD et al. 1999: 169
Lycodon albofuscus — LANZA 1999
Lepturophis borneensis — STUEBING & INGER 1999
Lepturophis albofuscus — HIEN et al. 2001
Lepturophis albofuscus — GRISMER et al. 2002
Lycodon albofuscus — DAS & YAAKOB 2007
Lycodon albofuscus — DAS 2012
Lepturophis albofuscus — WALLACH et al. 2014: 370
Lepturophis albofuscus — STUEBING et al. 2014
Lycodon albofuscus — FIGUEROA et al. 2016 
DistributionIndonesia (Sumatra, Nias); Thailand (Narathiwat, Satun, Songkhla, Surat Thani, Nakhon Si Thammarat, Yala),
Malaysia (Sabah, Sarawak, Borneo), Pulau Tioman.

Type locality: Sumatra

borneensis: Indonesia (Nias, Sumatra, Borneo);
Malaysia (Malaya and East Malaysia; Sabah, Sarawak, Selangor, Pahang, Jahore); S Thailand; Type locality: Sarawak. Emended to Kuching, Sarawak by de Rooij 1917).  
Reproductionoviparous. 
TypesHolotype: lost fide Iskandar & Colijn, 2001: 65. Not located at MNHN (I. Ineich, March 2019) 
DiagnosisAdditional details, e.g. a detailed description or comparisons (824 characters), are available for collaborators and contributors. Please contact us for details. 
CommentJuveniles have white bands on the black body while adults are completely black.

Type species: Sphecodes albo-fuscus DUMÉRIL, BIBRON & DUMÉRIL 1854: 394 is the type species of the genus Lepturophis BOULENGER 1900.

Synonymy: Dryocalamus is currently (2017) nested within Lycodon (e.g. see phylogenetic tree in Wostl et al. 2017), hence the Philippine Lycodon must be moved to a new genus or Lepturophis synonymized with Lycodon.

Habitat: fully arboreal (Harrington et al. 2018). 
References
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Boulenger, G.A. 1900. Description of new reptiles and batrachians from Bomeo. Proc. Zool. Soc. London: 182-187. - get paper here
  • Boulenger,G.A. 1885. A list of reptiles and batrachians from the Island of Nias. Ann. Mag. nat. Hist. (5) 16: 388-389 - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Cota, Michael; Robert W. Mendyk , Tanya Chan-ard , Sunchai Makchai and Pratheep Meewattana 2011. Range Extension, New Records and Natural History Notes on Lepturophis albofuscus (Duméril, Bibron & Duméril, 1854) (Squamata: Colubridae) in Thailand. The Thailand Natural History Museum Journal 5(2): 151-153
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • Das, I. & Yaakob, N. 2007. Status of knowledge of the Malaysian herpetofauna. In Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. in: L. S. L. Chua, L. G. Kirton & L. G. Saw (eds.), Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. Forest Research Institute Malaysia, Kepong, pp. 31-81
  • DAVID, Patrick; MEREL J. COX, OLIVIER S. G. PAUWELS, LAWAN CHANHOME AND KUMTHORN THIRAKHUPT 2004. Book Review - When a bookreview is not sufficient to say all: an in-depth analysis of a recent book on the snakes of Thailand, with an updated checklist of the snakes of the Kingdom. The Natural History Journal of Chulalongkorn University 4(1): 47-80 - get paper here
  • David,P. & Vogel, G. 1996. The snakes of Sumatra. An annotated checklist and key with natural history notes. Bücher Kreth, Frankfurt/M.
  • Duméril, A.M.C., G. BIBRON & A.H.A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 7 (partie 1). Paris, xvi + 780 S. - get paper here
  • Figueroa A, McKelvy AD, Grismer LL, Bell CD, Lailvaux SP 2016. A Species-Level Phylogeny of Extant Snakes with Description of a New Colubrid Subfamily and Genus. PLoS One 11 (9): e0161070.doi:10.1371/journal.pone.0161070 - get paper here
  • Fukuyama R, Fukuyama I, Kurita T, Kojima Y, Hossman MY, Noda A, Nishikawa K 2021. New herpetofaunal records from Gunung Mulu National Park and its surrounding areas in Borneo. Herpetozoa 34: 89-96 - get paper here
  • Grandison, A. G. C. 1978. Snakes of West Malaysia and Singapore. Annalen des Naturhistorischen Museums in Wien 81 [1977]: 283-303 - get paper here
  • Grandison, A.G.C. 1972. The Gunong Benom Expedition 1967. 5. Reptiles and amphibians of Gunong Benom with a description of a new species of Macrocalamus. Bull. Br. Mus. nat. Hist. (Zool.), London, 23: 45-101. - get paper here
  • GRISMER, L. LEE; EVAN S.H. QUAH, SHAHRUL ANUAR M.S,, MOHD ABDUL MUIN, PERRY L. WOOD, JR & SITI AZIZAH MOHD NOR 2014. A diminutive new species of cave-dwelling Wolf Snake (Colubridae: Lycodon Boie, 1826) from Peninsular Malaysia. Zootaxa 3815 (1): 051–067 - get paper here
  • Grismer, L.L. 2011. Amphibians and reptiles of the Seribuat Archipelago. Edition Chimaira, Frankfurt, 239 pp.
  • Grismer, L.L., McGuire, J.A.; Sosa, R. & Kaiser, H. 2002. Revised checklist and comments on the terrestrial herpetofauna of Pulau Tioman, Peninsular Malaysia. Herpetological Review 33 (1): 26-29 - get paper here
  • Grossmann, W. & Tillack, F. 2004. Pulau Tioman - Perle im Südchinesischen Meer, Teil 1. Reptilia (Münster) 9 (50): 42-49 - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Hien,P. Grossmann,W. & Schäfer, C. 2001. Beitrag zur Kenntnis der landbewohnenden Reptilienfauna von Pulau Tioman, West-Malaysia. Sauria 23 (4): 11-28 - get paper here
  • Janssen HY, Pham CT, Ngo HT, Le MD, Nguyen TQ, Ziegler T 2019. A new species of Lycodon Boie, 1826 (Serpentes, Colubridae) from northern Vietnam. ZooKeys 875: 1-29 - get paper here
  • Lanza,B. 1999. A new species of Lycodon from the Philippines, with a key to the genus (Reptilia: Serpentes: Colubridae). Tropical Zoology 12: 89-104 - get paper here
  • Lim, K.K.P. & Ng, H.H. 1999. The terrestrial herpetofauna of Pulau Tioman, Peninsular Malaysia. Raffles Bull. Zool., Suppl. No. 6: 131-155 - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. (ISBN 3-931587-12-6) - get paper here
  • Manthey,U. 1983. Exkursion am Mt. Kinabalu (4101 m), Nordborneo, Teil 3: Checkliste der Herpetofauna oberhalb 600 m ü. NN. Herpetofauna 5 (23): 20-31 - get paper here
  • Murphy, J. C.; Voris, H. K.; Karns, D. R. 1994. A field guide and key to the snakes of the Danum Valley, A bornean tropical rainforest ecosystem. Bull. Chicago Herp. Soc. 29: 133-151
  • Onn, Chan Kin; Mohd. Shahfiz Azman, Nor Azlin and Pan Khang Aun 2009. Additions to the Herpetofauna of Pasoh Forest Reserve, Negeri Sembilan, Peninsular Malaysia. Tropical Life Sciences Research, 20 (1): 71–80 - get paper here
  • Rooijen, J. van & van Rooijen, M. 2002. Einige Ergänzungen, Berichtigungen und neue Beobachtungen zur Herpetofauna von Pulau Tioman, West-Malaysia. Sauria 24 (3): 3-12 [erratum in 24 (4): 34] - get paper here
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  • Stuebing, R.B. & INGER, R.F. 1999. A field guide to the snakes of Borneo. Natural history Publications (Borneo), Kota Kinabalu, 254 pp. [corrections in HR 31: 124]. - get paper here
  • Stuebing, R.B., Inger, R.F. & Lardner, B. 2014. A field guide to the snakes of Borneo, second edition. Natural history Publications (Borneo)
  • Syafiq MF, Badli-Sham BH, Grismer LL, Ahmad AB 2023. Uneven species occurrence and richness of lowland snakes (Serpentes, Squamata) in Terengganu, Peninsular Malaysia, with new locality records. ZooKeys 1168: 11–39
  • Syafiq, Muhamad Fatihah; Baizul Hafsyam Badli-Sham, Noor-Shahirah Ibrahim, Lukman H. B. Ismail, Muhammad Afif Mohd Amin, Goh Shu Xian, Rabiatul ‘Aqilah Mohamad Ariffin, Muhammad Afiq-Suhaimi, Lok Kar Men, Dila Adilina Danelo, Mohamad Aqmal-Naser, Muh 2024. TAXONOMIC COMPOSITION, DIVERSITY, AND CONSERVATION STATUS OF REPTILIAN FAUNA AT SEKAYU LOWLAND FORESTS, TERENGGANU, PENINSULAR MALAYSIA. Russian Journal of Herpetology Vol. 31, No. 1, 2024, pp. 14 – 23 - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Vogel, G. & R. Kreutz 1990. Erstnachweis von Lepturophis borneensis BOULENGER 1900 für Thailand. Sauria 12 (1): 21-23 - get paper here
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  • WOSTL, ELIJAH; AMIR HAMIDY, NIA KURNIAWAN, ERIC N. SMITH 2017. A new species of Wolf Snake of the genus Lycodon H. Boie in Fitzinger (Squamata: Colubridae) from the Aceh Province of northern Sumatra, Indonesia. Zootaxa 4276 (4): 539–553 - get paper here
 
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