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Lycodon futsingensis (POPE, 1928)

IUCN Red List - Lycodon futsingensis - Least Concern, LC

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Common Names 
SynonymDinodon futsingensis POPE 1928
Dinodon septentrionale ruhstrati — MAKI 1931 (part.)
Dinodon futsingensis — BOURRET 1934
Dinodon futsingense — BOURRET 1936
Lycodon ruhstrati — POPE 1935 (part.)
Lycodon ruhstrati ruhstrati — OTA 1988 (part.)
Lycodon ruhstrati ruhstrati — LANZA 1999 (part.)
Ophites ruhstrati — ZHAO & ADLER 1993 (part.)
Lycodon futsingensis — VOGEL et al. 2009
Lycodon futsingensis — GUO et al. 2013
Lycodon futsingensis — WALLACH et al. 2014: 394 
DistributionChina (Fukien = Fujian, Hong Kong, Guangdong)
Vietnam (Vin Puh, Bắc Kạn [Ngan-Son], Lào Cai, Hà Tĩnh, Quang Binh (Phong Nha-Ke Bang National Park), Laos

Type locality. “Futsing Hsien, Fukien Province”, now Fuqing Xian [County], 25°44’N, 119°22’E, Fujian Province, People’s Republic of China.  
TypesHolotype: AMNH 34106, collected by C.H. Pope, August-October, 1925. – Paratype. AMNH R34105, same data as the holotype. 
DiagnosisDiagnosis. A species of the genus Lycodon, characterized by: (1) a loreal not entering orbit; (2) 17 dorsal scale rows at the forepart of the body and at midbody; (3) the upper dorsal and vertebral scale rows entirely smooth; (4) 193-204 ventrals in males and 198-208 in females; (5) 72-87 SC in males and 78-85 in females; (6) a relative tail length ratio of 0.194-0.229 in males and 0.194-0.216 in females; (7) 19-33 light bands on a dark body; (8) the first band starting at Ve 13-21; and (9) upper maxillary teeth: 7-8 + 3-4 + 2-3. This species can be recognized by the combination of unkeeled dorsal scale rows and low ventral and subcaudal counts. For a detailed comparisons with other species of Lycodon see discussion in VOGEL eta l. (2009). 
CommentSynonymy: the species has been synonymized with Lycodon ruhstrati (Pope, 1929), to which it is morphologically similar (Pope, 1935; Vogel et al., 2009). This arrangement has been followed for a long time by some authors (Zhao et al., 1998; Lanza, 1999). Recently, on the basis of morphological comparison, Vogel et al. (2009) revalidated Dinodon futsingensis and recognized it as Lycodon futsingensis. 
EtymologyEtymology. This species was named according to the name of the type locality, Futsing Hsien, now spelt as Fuxing Qian in Pinyin. 
  • Bourret, R. 1934. Notes herpétologiques sur l'Indochine Française II. Sur quelques serpents des montagnes du Tonkin. Bull. Gen. Instr. Pub. Hanoi 8 (April 1934): 149-157 [13: 1-11 ?]
  • Bourret, R. 1936. Les Serpents de l'Indochine, vol. 2. Imprimerie Henri Basuyau & Cie, Toulouse.
  • Deuve, J. 1961. Liste annotee des Serpents du Laos. Bull. Soc. Sci. Nat. Laos 1:5-32.
  • Guo, Peng; Liang Zhang, Qin Liu, Cao Li, R. Alexander Pyron, Ke Jiang, Frank T. Burbrink 2013. Lycodon and Dinodon: One genus or two? Evidence from molecular phylogenetics and morphological comparisons. Molecular Phylogenetics and Evolution 68 (1): 144–149 - get paper here
  • Hecht, Vera L.; Cuong T. Pham, Tao T. Nguyen, Truong Q. Nguyen, Michael Bonkowski & Thomas Ziegler 2013. First report on the herpetofauna of Tay Yen Tu Nature Reserve, northeastern Vietnam. Biodiversity Journal 4 (4): 507–552
  • Lanza,B. 1999. A new species of Lycodon from the Philippines, with a key to the genus (Reptilia: Serpentes: Colubridae). Tropical Zoology 12: 89-104 - get paper here
  • Luu, Vinh Quang; Michael Bonkowski, Truong Quang Nguyen, Minh Duc Le,, Thomas Calame & Thomas Ziegler 2018. A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from central Laos. Revue suisse de Zoologie 125(2): 263-276
  • Maki, M. 1931. Monograph of the Snakes of Japan. Dai-ichi Shobo, Tokyo. (1), 7, 240 pages (expanded edition 1933)
  • NEANG, THY; TIMO HARTMANN, SEIHA HUN, NICHOLAS J. SOUTER & NEIL M. FUREY 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068–080
  • Nguyen, Luan Thanh; Ha Van Hoang, Thang Tai Nguyen, McCormack Timothy E.M., Sang Ngoc Nguyen 2016. A COLLECTION OF AMPHIBIANS AND REPTILES FROM BAC HUONG HOA NATURE RESERVE, QUANG TRI PROVINCE, VIETNAM - get paper here
  • Nguyen, Truong Quang 2017. New records of amphibians and reptiles from Ha Giang Province, Vietnam Herpetology Notes 10: 183-191 - get paper here
  • Ota, Hidetoshi 1988. Taxonomic notes on Lycodon ruhstrati (Colubridae: Ophidia) from East Asia. Journal of Taiwan Museum 41 (1): 85-91
  • Pope, CLIFFORD H. 1928. Seven new reptiles from Fukien Province, China. American Museum Novitates 320: 1-6. - get paper here
  • Pope, CLIFFORD H. 1935. The Reptiles of China. Turtes, Crocodilians, Snakes, Lizards. Amer. Mus. Nat. Hist., New York, Nat. Hist. Central Asia, 10: lii, 1-604
  • Vogel, G. & Luo, J. 2011. A new species of the genus Lycodon (Boie, 1826) from the southwestern mountains of China (Squamata: Colubridae). Zootaxa 2807: 29–40 - get paper here
  • Vogel, G., T. Q. Nguyen, P. Kingsada & T. Ziegler 2012. A new species of the genus Lycodon BOIE, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology, 8: 344-352 - get paper here
  • Vogel, G.; David, P.; Pauwels, O.S.G.; Sumontha, M.; Norval, G.; Hendrix, R.; Vu, N.T. & Ziegler, T. 2009. A revision of Lycodon ruhstrati (Fischer 1886) auctorum (Squamata Colubridae), with the description of a new species from Thailand and a new subspecies from the Asian mainland. Tropical Zoology 22: 131-182 - get paper here
  • Vogel, Gernot 2010. Zur Systematik der Schlangen der orientalischen Region: Forschungsdefizite führen zur Unterschätzung der Artenvielfalt. Ophidia 4 (2): 18-26 - get paper here
  • Vogel, Gernot & Patrick David 2010. A new species of the genus Lycodon (Boie, 1826) from Yunnan Province, China (Serpentes: Colubridae). Bonn Zool. Bull. 57 (2): 289–296 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • ZHANG Liang; JIANG Ke, HU Ping, YU Bao-Cheng, PENG Bo-Yong, TANG Xiao-Ping, HU Hui-Jian 2011. Lycodon futsingensis: A New Snake Record in Guangdong Province,China. Chinese Journal of Zoology 46 (1):128-130
  • Zhang, Liang and Kai Wang. 2014. Lycodon futsingensis (Fuqing wolf snake) diet. Herpetological Review 45 (4): 711-712
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
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