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Lycodon ruhstrati (FISCHER, 1886)

IUCN Red List - Lycodon ruhstrati - Least Concern, LC

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
SubspeciesLycodon ruhstrati abditus VOGEL, DAVID, PAUWELS, SUMONTHA, NORVAL, HENDRIX, RU & ZIEGLER 2009
Lycodon ruhstrati ruhstrati (FISCHER 1886) 
Common NamesE: Mountain Wolf Snake, Formosa Wolf Snake
G: Berg-Wolfszahnnatter 
SynonymOphites Ruhstrati FISCHER 1886: 16
Dinodon septentrionalis — BOULENGER 1893: 363 (part.)
Dinodon septentrionalis var. ruhstrati — BOULENGER 1899: 165
Dinodon septentrionale ruhstrati — STEJNEGER 1907: 370
Ophites ruhstrati — ZHAO & ADLER 1993
Lycodon ruhstrati — LANZA 1999
Dinodon ruhstrati — WEB SITE
Lycodon ruhstrati — ZIEGLER 2002: 241
Lycodon cf. ruhstrati — ZIEGLER et al. 2007
Lycodon ruhstrati — WALLACH et al. 2014: 395 
DistributionTaiwan, China (Jiangxi, Fujian, Guangdong, northward to Anhui and west to Sichuan [elevation 800-1850 m] and SE Gansu), N Vietnam (elevation 500-1500 m)

ruhstrati: Taiwan (endemic)

abditus: Type locality: U Bo region, Phong Nha – Ke Bang NP, Quang Binh Province, Vietnam.

Type locality: Süd-Formosa [= South Taiwan], China. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
TypesSyntypes: LMNMO 918 (formerly LMNMO 37c & 73), a 583+ mm male, and LMNMO 919 (formerly LMNMO 37b & 74)
Syntypes. REP 918-19
Holotype (abditus): ZFMK 86451 (GenBank: EU999209), adult female. Collected by Thomas Ziegler, 20 June 2006 (mentioned as Lycodon cf. ruhstrati in ZieGler et al. 2007). 
CommentTaxonomy and nomenclature: For a discussion of nomenclatorial issues concerning the genera Lycodon and Ophites see Zhao & Adler (who listed this species under Ophites). Dinodon futsingensis POPE 1928, previously a synonym of L. ruhstrati, has been revalidated by VOGEL et al. 2009. Lycodon ruhstrati multifasciatus (MAKI 1931) has been elevated to full species status by VOGEL et al. 2009. 
EtymologyNamed after Mr. Ruhstrat, who collected the types in southern Taiwan.

abditus: The subspecific nomen of this subspecies is the Latin adjective abditus (-a, -um), meaning “hidden”, in allusion to the long confusion of this species with Lycodon ruhstrati ruhstrati. 
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Boulenger, GEORGE A. 1899. On a collection of reptiles and batrachians made by Mr. J. D. La Touche in N.W. Fokien, China. Proc. Zool. Soc. London 1899: 159-172 - get paper here
  • Fischer, J. G. 1886. Herpetologische Notizen. Abh. Geb. Naturw. Hamb. 9: 51-67 (1-19) - get paper here
  • KUCHARZEWSKI, CH. & TILLACK, F. & FUHRMANN, K. 2009. Rediscovery and redescription of the type specimens of Ophites ruhstrati FISCHER, 1886. Herpetozoa 22: - get paper here
  • Lanza,B. 1999. A new species of Lycodon from the Philippines, with a key to the genus (Reptilia: Serpentes: Colubridae). Tropical Zoology 12: 89-104 - get paper here
  • Mori,M. 1982. Japans Schlangen, Vols. 1-3 (80, 102, 123 pp.). Tokyo (Igaku-Shoin Ltd.) [bilingual edition in German and Japanese]
  • NEANG, THY; TIMO HARTMANN, SEIHA HUN, NICHOLAS J. SOUTER & NEIL M. FUREY 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068–080
  • NORVAL, G. & J.-J. MAO 2006. Die Berg-Wolfszahnnatter (Lycodon ruhstrati ruhstrati [FISCHER, 1886]): Eine Beschreibung ihres Abwehrverhaltens. Sauria 28 (4): 47-50 - get paper here
  • Norval, G. & Mao, J.J. 2008. An instance of Japalura swinhonis as prey of Lycodon ruhstrati ruhstrati in Chiayi County, Taiwan. Sauria 30 (3): 43-45 - get paper here
  • Norval, G. & Mao, J.J. 2008. An instance of arboricolous predation by a Mountain Wolf Snake (Lycodon ruhstrati ruhstrati [FISCHER], 1886]) on a Brown anole (Norops sagrei DUMÉRIL & BIBRON, 1837). Sauria 30 (1): 59-62 - get paper here
  • Norval, Gerrut; Charles R. Bursey, Stephen R. Goldberg and Jean-Jay Mao. 2009. The first record of Kiricephalus pattoni Stephens, 1908 as a parasite of Lycodon ruhstrati ruhstrati Fischer, 1886 from Chiayi County, Taiwan. Herpetology Notes 2: 79-80. - get paper here
  • Orlov, N. L., R. W. Murphy & T. J. Papenfuss 2000. List of snakes of Tam-Dao mountain ridge (Tonkin, Vietnam). Russ. J. Herpetol. 7 (1): 69-80. - get paper here
  • Ota, H. 2000. Current status of the threatened amphibians and reptiles of Japan. Popul. Ecol. 42: 5-9
  • Ota, Hidetoshi 1988. Taxonomic notes on Lycodon ruhstrati (Colubridae: Ophidia) from East Asia. Journal of Taiwan Museum 41 (1): 85-91
  • Pope, CLIFFORD H. 1928. Seven new reptiles from Fukien Province, China. American Museum Novitates 320: 1-6. - get paper here
  • Sang, Nguyen Van; Ho Thu Cuc, Nguyen, Quang Truong 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Stejneger, L. 1910. The batrachians and reptiles of Formosa. Proc. US Natl. Mus. 38: 91-114 - get paper here
  • Stejneger, LEONHARD H. 1907. Herpetology of Japan and adjacent territory. Bull. US Natl. Mus. 58: xx, 1-577 - get paper here
  • Vogel, G. & Brachtel, N. 2008. Contribution to the knowledge of Lycodon ruhstrati (FISCHER, 1886) in Vietnam - taxonomy and biology of a little-known species. Salamandra 44 (4): 207-224 - get paper here
  • Vogel, G. & Luo, J. 2011. A new species of the genus Lycodon (Boie, 1826) from the southwestern mountains of China (Squamata: Colubridae). Zootaxa 2807: 29–40 - get paper here
  • Vogel, G., T. Q. Nguyen, P. Kingsada & T. Ziegler 2012. A new species of the genus Lycodon BOIE, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology, 8: 344-352 - get paper here
  • Vogel, G.; David, P.; Pauwels, O.S.G.; Sumontha, M.; Norval, G.; Hendrix, R.; Vu, N.T. & Ziegler, T. 2009. A revision of Lycodon ruhstrati (Fischer 1886) auctorum (Squamata Colubridae), with the description of a new species from Thailand and a new subspecies from the Asian mainland. Tropical Zoology 22: 131-182 - get paper here
  • Vogel, Gernot 2010. Zur Systematik der Schlangen der orientalischen Region: Forschungsdefizite führen zur Unterschätzung der Artenvielfalt. Ophidia 4 (2): 18-26 - get paper here
  • Vogel, Gernot & Patrick David 2010. A new species of the genus Lycodon (Boie, 1826) from Yunnan Province, China (Serpentes: Colubridae). Bonn Zool. Bull. 57 (2): 289–296 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
  • Zhao, E. (ed.) 2003. Coloured atlas of Sichuan reptiles [in Chinese]. Beijing, China Forestry Publishing House
  • Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp.
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
  • Ziegler, T. 2002. Die Amphibien und Reptilien eines Tieflandfeuchtwald-Schutzgebietes in Vietnam. Natur und Tier Verlag (Münster), 342 pp. - get paper here
  • ZIEGLER, THOMAS; RALF HENDRIX, VU NGOC THANH, MARTINA VOGT, BERNHARD FORSTER & DANG NGOC KIEN 2007. The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key. Zootaxa 1493: 1-40 - get paper here
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