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Lygodactylus angularis GÜNTHER, 1893

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
SubspeciesLygodactylus angularis angularis GÜNTHER 1893
Lygodactylus angularis grzimeki BANNIKOV & DAREVSKY 1969 
Common NamesE: Angulated dwarf gecko 
SynonymLygodactylus angularis GÜNTHER 1893: 555
Lygodactylus angularis angularis — LOVERIDGE 1953: 169
Lygodactylus angularis — KLUGE 1993
Lygodactylus angularis angularis — BROADLEY & HOWELL 1991: 9
Lygodactylus (Lygodactylus) angularis angularis — RÖSLER 2000: 92
Lygodactylus angularis —RÖLL et al. 2010
Lygodactylus angularis — SPAWLS et al. 2018: 101

Lygodactylus angularis grzimeki BANNIKOV & DAREVSKY 1969
Lygodactylus angularis grzimeki — BROADLEY & HOWELL 1991: 9
Lygodactylus (Lygodactylus) angularis grzimeki — RÖSLER 2000: 92
Lygodactylus grzimeki — RÖLL et al. 2010 
DistributionSW Tanzania, Kenya, Zimbabwe (Shire Highlands, Nyassaland), Malawi, Mozambique, Zambia, Zaire (Katanga, Kapiri) = Democratic Republic of the Congo

angularis: SW Tanzania, Malawi, Mozambique, Zambia. Type locality: Shire Highlands, Malawi.

heeneni: S Zaire, Zambia

grzimeki: Lake Manyara, Tanzania. Type locality: Lake Manyara National Park, Tanzania.  
Reproductionoviparous 
TypesType: BMNH 1946.8.22.57 (and possibly additional specimens).
Holotype: ZISP 18097 = ZIN (Zoological Institute of the Academy of Sciences of the USSR, Leningrad), female [grzimeki] 
Diagnosis 
CommentDistribution: known in Kenya only from a single specimen from Mombasa (CAS).

Subspecies: Lygodactylus angularis heeneni is now considered as a valid species. 
EtymologyL. a. heeneni was named after M. HEENEN, Gouvernor of the Province of Katanga.

L. a. grzimeki was named after Bernhard Grzimek (1909–1987), a renowned German zoo director, zoologist and animal conservationist 
References
  • Bannikov, A. G., and l. S. Darevsky. 1969. Lygodactylus angularis grzimeki subsp. n. Bannikov et Darevsky from the Lake Manyara National Park, Tanzania [in Russian]. Zoologicheskii Zhurnal, 48:452—455.
  • Bedriaga, J. von 1886. Beiträge zur Kenntnis der Lacertiden-Familie (Lacerta, Algiroides, Tropidosaura, Zerzumia, Bettaia). Abh. senckenb. naturf. Ges. (Frankfurt) 14: 17-444 - get paper here
  • Branch, W.R.; Rödel, M.-O. & Marais, J. 2005. Herpetological survey of the Niassa Game Reserve, northern Mozambique - Part I: Reptiles. Salamandra 41 (4): 195-214 - get paper here
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Broadley, D.G. 1991. The Herpetofauna of Northern Mwinilunga Distr., Northw. Zambia. Arnoldia Zimbabwe 9 (37): 519-538
  • Broadley, D.G. 1998. The reptilian fauna of the Democratic Republic of the Congo (Congo-Kinshasa). In: Schmidt, K.P. and Noble, G.K., Contributions to the Herpetology of the Belgian Congo... [reprint of the 1919 and 1923 papers]. SSAR Facsimile reprints in Herpetology, 780 pp.
  • Buruwate, T. C., & Lloyd-Jones, D. J. 2024. Amphibian and Reptile Diversity of Niassa Special Reserve, Northern Mozambique. Journal of East African Natural History, 113(1), 1-18 - get paper here
  • Castiglia, Riccardo and Flavia Annesi 2011. The phylogenetic position of Lygodactylus angularis and the utility of using the 16S rDNA gene for delimiting species in Lygodactylus (Squamata, Gekkonidae). Acta Herpetologica 6 (1): 35-45 - get paper here
  • Günther,A. 1893. Report on a collection of reptiles and batrachians transmitted by Mr. H. H. Johnston, C. B., from Nyassaland. Proc. Zool. Soc. London 1892: 555-558 [1892] - get paper here
  • Haagner,G.V.; Branch,W.R. & Haagner,A.J.F. 2000. Notes on a collection of reptiles from Zambia and adjacent areas of the Democratic Republic of the Congo. Annals of the Eastern Cape Museum 1: 1 – 25
  • Lobón-Rovira, J., Bauer, A. M., Vaz Pinto, P., Trape, J. F., Conradie, W., Kusamba, C., ... & Greenbaum, E. 2023. Integrative revision of the Lygodactylus gutturalis (Bocage, 1873) complex unveils extensive cryptic diversity and traces its evolutionary history. Zoological Journal of the Linnean Society, zlad123 - get paper here
  • Loveridge, A. 1953. Zoological Results of a fifth expedition to East Africa. III. Reptiles from Nyasaland and Tete. Bull. Mus. Comp. Zool. Harvard 110 (3): 142-322. - get paper here
  • Pietersen, Darren, Verburgt, Luke & Davies, John 2021. Snakes and other reptiles of Zambia and Malawi. Struik Nature / Penguin Random House South Africa, 376 pp., ISBN 9781775847373
  • PORTIK, DANIEL M.; SCOTT L. TRAVERS, AARON M. BAUER, WILLIAM R. BRANCH 2013. A new species of Lygodactylus (Squamata: Gekkonidae) endemic to Mount Namuli, an isolated ‘sky island’ of northern Mozambique. Zootaxa 3710 (5): 415–435 - get paper here
  • Röll, B. 2018. Tagaktive, kleine Geckos – die Gattung Lygodactylus. Reptilia 23 (132): 16-23
  • Röll, Beate; Heike Pröhl, Klaus-Peter Hoffmann 2010. Multigene phylogenetic analysis of Lygodactylus dwarf geckos (Squamata: Gekkonidae). Molecular Phylogenetics and Evolution 56 (1): 327-335 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Spawls, S. & Rotich, D. 1997. An annotated checklist of the lizards of Kenya. J. East African Nat. Hist. 86: 61-83
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Witte, G. F. de 1933. Description de Reptiles nouveaux provenant du Katanga (1930-31). Rev. Zool. Bot. afr., Bruxelles, 23 (2): 185-192
  • Witte, G. F. de 1933. Reptiles récoltés au Conge Belge par le Dr. H. Schouteden et par M. G.-F. Witte. Ann. Mus. Conge belge Zool. Ser. 1 Tome III: 53-100.
  • Witte, G. F. de 1953. Reptiles. Exploration du Parc National de l'Upemba. Mission G. F. de Witte en collaboration avec W. Adam, A. Janssens, L. Van Meel et R. Verheyen (1946–1949). Institut des Parcs Nationaux du Congo Belge. Brussels, vol. 6, 322 pp. - get paper here
 
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