Lygosoma angeli (SMITH, 1937)

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Higher Taxa	Scincidae, Lygosominae (Lygosomini), Scincoidea, Sauria, Squamata (lizards)
Subspecies
Common Names	E: Angel's Writhing Skink
Synonym	Riopa angeli SMITH 1937: 366 Lygosoma angeli — GREER 1977 Lygosoma angeli — BOBROV 1995 Lygosoma angeli — ZIEGLER et al. 2007 Riopa angeli — BOBROV & SEMENOV 2008 Lygosoma angeli — COTA et al. 2011 Lygosoma angeli — STUART et al. in WCS 2024
Distribution	Vietnam, Thailand, Laos, E Cambodia Type locality: Trang Bom, Bien Hoa (Dong Nai Province, S Vietnam)
Reproduction	oviparous (not imputed, fide Zimin et al. 2022)
Types	Holotype: MNHN-RA 1937.0021 (by original designation), collected by Mlle Colani.
Diagnosis	Diagnosis: The species differs from its congeners by the combination of the following characters: SVL 76– 112 mm, equal to TaL; TrunkL 3.4–4.2 times longer than SFlL; lower eyelid scaly; supranasals in contact; frontal as long as frontoparietal; frontoparietal single; nuchals absent; external ear opening with a small lobule; supralabials 7; dorsal scales smooth (see Fig. 9A); scale rows around midbody 30; paravertebral scales 107–115; ventral scales 112–123; limbs very short (SVL/FlL: about 20; SVL/HlL: about 13); subdigital lamellae under fourth toe 6–7 (Smith 1937b; pers. observ.) (Geissler et al. 2011).
Comment
References	Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here Bobrov V.V., Semenov D.V. 2008. Lizards of Vietnam [in Russian]. Moscow, 236 pp. Bobrov, V. V. 1995. Checklist and bibliography of lizards of Vietnam. Smithsonian Herp. Inf. Serv. (105): - get paper here Cota, Michael; Peter Geissler , Tanya Chan-ard and Sunchai Makchai 2011. First Record of Lygosoma angeli (Smith, 1937) (Reptilia: Squamata: Scincidae) in Thailand with Notes on Other Specimens from Laos. The Thailand Natural History Museum Journal 5(2): 125-132 GEISSLER, PETER; TIMO HARTMANN & THY NEANG 2012. A new species of the genus Lygosoma Hardwicke & Gray, 1827 (Squamata: Scincidae) from northeastern Cambodia, with an updated identification key to the genus Lygosoma in mainland Southeast Asia. Zootaxa 3190: 56–68 - get paper here Geissler, Peter; Truong Quang Nguyen, Trung My Phung, Robert Wayne Van Devender, Timo Hartmann, Balázs Farkas, Thomas Ziegler and Wolfgang Böhme 2011. A review of Indochinese skinks of the genus Lygosoma Hardwicke & Gray, 1827 (Squamata: Scincidae), with natural history notes and an identification key. Biologia 66 (6): 1159-1176 - get paper here Greer, A.E. 1977. The systematics and evolutionary relationships of the scincid lizard genus Lygosoma. Journal of Natural History 11: 515-540. - get paper here Guibé, J. & Davis, D.D. 1951. Fernand Angel, 1881-1950. Copeia 1951 (1): 1-2 - get paper here Neang, Thy; Menghor Nut 2015. First record of Lygosoma angeli (Smith, 1937) (Squamata: Scincidae) from eastern Cambodia. Herpetology Notes 8: 321-322 - get paper here Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp. Smith, M. A. 1937. Un nouveau Lézard de Cochinchine. Bulletin du Muséum National d’Histoire Naturelle, (2) 9(6): 366 - get paper here Smith, M.A. 1937. A review of the genus Lygosoma (Scincidae: Reptilia) and its allies. Records of the Indian Museum 39 (3): 213-234 WCS 2024. Wildlife in Lao PDR, 2024 Status Report. Vientiane; Wildlife Conservation Society Lao PDR Program Ziegler, T; Schmitz, A; Heidrich, A; Vu, NT; Nguyen, QT 2007. A new species of Lygosoma (Squamata: Sauria: Scincidae) from the Central Truong Son, Vietnam, with notes on its molecular phylogenetic position. Revue Suisse de Zoologie 114 (2): 397-415 - get paper here Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
External links	IUCN Red List http://novataxa.blogspot.com/2012/02/2011-indochinese-lygosoma.html http://www.wcmc.org.uk/infoserv/countryp/vietnam/app2.html Google images

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