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Lygosoma angeli (SMITH, 1937)

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Higher TaxaScincidae, Lygosominae (Lygosomini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Angel's Writhing Skink 
SynonymRiopa angeli SMITH 1937: 366
Lygosoma angeli — GREER 1977
Lygosoma angeli — BOBROV 1995
Lygosoma angeli — ZIEGLER et al. 2007
Riopa angeli — BOBROV & SEMENOV 2008
Lygosoma angeli — COTA et al. 2011 
DistributionVietnam, Thailand, Laos, E Cambodia

Type locality: Trang Bom, Bien Hoa (Dong Nai Province, S Vietnam)  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: MNHN-RA 1937.0021 (by original designation), collected by Mlle Colani. 
DiagnosisDiagnosis: The species differs from its congeners by the combination of the following characters: SVL 76– 112 mm, equal to TaL; TrunkL 3.4–4.2 times longer than SFlL; lower eyelid scaly; supranasals in contact; frontal as long as frontoparietal; frontoparietal single; nuchals absent; external ear opening with a small lobule; supralabials 7; dorsal scales smooth (see Fig. 9A); scale rows around midbody 30; paravertebral scales 107–115; ventral scales 112–123; limbs very short (SVL/FlL: about 20; SVL/HlL: about 13); subdigital lamellae under fourth toe 6–7 (Smith 1937b; pers. observ.) (Geissler et al. 2011). 
Comment 
EtymologyNamed after Fernand Angel (1881-1950), French herpetologist. See Guibé & Davis (1951) for an obituary. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bobrov V.V., Semenov D.V. 2008. Lizards of Vietnam [in Russian]. Moscow, 236 pp.
  • Bobrov, V. V. 1995. Checklist and bibliography of lizards of Vietnam. Smithsonian Herp. Inf. Serv. (105): - get paper here
  • Cota, Michael; Peter Geissler , Tanya Chan-ard and Sunchai Makchai 2011. First Record of Lygosoma angeli (Smith, 1937) (Reptilia: Squamata: Scincidae) in Thailand with Notes on Other Specimens from Laos. The Thailand Natural History Museum Journal 5(2): 125-132
  • GEISSLER, PETER; TIMO HARTMANN & THY NEANG 2012. A new species of the genus Lygosoma Hardwicke & Gray, 1827 (Squamata: Scincidae) from northeastern Cambodia, with an updated identification key to the genus Lygosoma in mainland Southeast Asia. Zootaxa 3190: 56–68 - get paper here
  • Geissler, Peter; Truong Quang Nguyen, Trung My Phung, Robert Wayne Van Devender, Timo Hartmann, Balázs Farkas, Thomas Ziegler and Wolfgang Böhme 2011. A review of Indochinese skinks of the genus Lygosoma Hardwicke & Gray, 1827 (Squamata: Scincidae), with natural history notes and an identification key. Biologia 66 (6): 1159-1176 - get paper here
  • Greer, A.E. 1977. The systematics and evolutionary relationships of the scincid lizard genus Lygosoma. Journal of Natural History 11: 515-540. - get paper here
  • Guibé, J. & Davis, D.D. 1951. Fernand Angel, 1881-1950. Copeia 1951 (1): 1-2 - get paper here
  • Neang, Thy; Menghor Nut 2015. First record of Lygosoma angeli (Smith, 1937) (Squamata: Scincidae) from eastern Cambodia. Herpetology Notes 8: 321-322 - get paper here
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Smith, M. A. 1937. Un nouveau Lézard de Cochinchine. Bulletin du Muséum National d’Histoire Naturelle, (2) 9(6): 366 - get paper here
  • Smith, M.A. 1937. A review of the genus Lygosoma (Scincidae: Reptilia) and its allies. Records of the Indian Museum 39 (3): 213-234
  • Ziegler, T; Schmitz, A; Heidrich, A; Vu, NT; Nguyen, QT 2007. A new species of Lygosoma (Squamata: Sauria: Scincidae) from the Central Truong Son, Vietnam, with notes on its molecular phylogenetic position. Revue Suisse de Zoologie 114 (2): 397-415 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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