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Lygosoma kinabatanganensis GRISMER, QUAH, DUZULKAFLY & YAMBUN, 2018

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Higher TaxaScincidae, Lygosominae, Scincoidea, Sauria, Squamata (lizards) 
Subspecies 
Common NamesKinabatangan Supple Skink 
SynonymLygosoma kinabatanganensis GRISMER, QUAH, DUZULKAFLY & YAMBUN 2018
Riopa bampfyldei — MALKMUS et al. 1999: 195
Riopa bampfyldei — MALKMUS et al., 2002: 281
Lygosoma bampfyldei — DAS 2010: 238
Lygosoma bampfyldei — KARIN et al. 2018: 246 
DistributionEast Malaysia (Sabah)

Type locality: east coast residency, Kinabatangan District, Deramakot camp (now the Deramakot Sabah Forestry Department), Sabah, East Malaysia (5°17N; 117°33E; 146 m).  
Reproduction 
TypesHolotype: FMNH 76226, Adult male, collected by Robert F. Inger on 10 May 1956. 
DiagnosisDiagnosis. Lygosoma kinabatanganensis sp. nov. can be differentiated from all other Lygosoma by having the combination of a large (SVL = 141 mm) robust body (PEC/SVL = 0.15); seven supralabials; six or seven infralabials; midline contact of the supranasals; prefrontals not in contact; frontoparietal contacting four supraoculars; eight superciliaries; three postsuboculars, the first being large; one primary and three secondary and tertiary temporals; four nuchal scales; a deep postnasal groove extending from the nasal scale to below the anterior portion of the eye and lying below the anterior loreals and lower preocular and above the second and third supralabials; scaly lower eyelid, no window; 42 midbody scale rows; 98 paravertebral scale rows; 104 ventral scale rows; 39 caudal scale rows at the tenth subcaudal; nine relatively small precloacal scales; smooth to weakly keeled subdigital finger lamellae, 10 lamellae on third finger; keeled subdigital toe lamellae, 15 lamellae on fourth toe; low, round, small palmar scales numbering eight across the base of the palm; head pattern consisting of a dark, frontal and occipital bands separated by a yellowish band; dark occipital band not confluent with lighter color of dorsum and tail; sides of tail olive. These characters are scored across all species in the L. bampfyldei group in Table 2 (Grismer et al. 2018).

Comparisons. Lygosoma kinabatangensis sp. nov. differs from all other species in the bampfyldei group by the frontoparietal contacting five vs. four supraoculars, having 104 vs. 94–99 ventral scales, 39 vs. 28–34 scales around the tail, and nine relatively small vs. seven or eight large precloacal scales. Other characters differentially separating L. kinabatanganensis sp. nov. from various combinations of the other species are listed in Table 1 (Grismer et al. 2018).
 
Comment 
EtymologyThe specific epithet, kinabatanganensis is in reference to this species being from the Kinabatangan region of Sabah in East Malaysia. 
References
  • Das, I. 2010. A Field Guide to the Reptiles of Southeast Asia. New Holland, London. 376 pp. [book review in Sauria 33 (3): 51, and Russ. J. Herp. 18: 325] - get paper here
  • GRISMER, L. LEE; EVAN S. H. QUAH, ZAHARIL DUZULKAFLY & PAUL YAMBUN 2018. On the taxonomy of Lygosoma bampfyldei Bartlett, 1895 (Squamata: Scincidae) with descriptions of new species from Borneo and Peninsular Malaysia and the resurrection of Lygosoma schneideri Werner, 1900. Zootaxa 4438 (3): 528–550 - get paper here
  • KARIN, BENJAMIN R.; ELYSE S. FREITAS, SAMUEL SHONLEBEN, L. LEE GRISMER, AARON M. BAUER, INDRANEIL DAS 2018. Unrealized diversity in an urban rainforest: A new species of Lygosoma (Squamata: Scincidae) from western Sarawak, Malaysia (Borneo) Zootaxa 4370 (4): 345-362 - get paper here
  • Malkmus, R., Brühl, C. & Eltz, T. 1999. Amfibieën en reptielen Deramakot (Sabah, Maleisië). Lacerta, 57, 191–198
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
 
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