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Macropholidus montanuccii TORRES-CARVAJAL, VENEGAS & NUNES, 2020

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Common NamesE: Montanucci’s Cuilanes.
Spanish: Cuilanes de Montanucci 
SynonymMacropholidus montanuccii TORRES-CARVAJAL, VENEGAS & NUNES 2020
Macropholidus labiopunctatus — PARRA et al. 2020 (nomen nudum) 
DistributionPeru (Piura)

Type locality: Peru, Departamento de Piura, Provincia Ayabaca, Bosque de Cuyas, 4°35′45.9′′S, 79°42′46.6′′W, WGS84, 2,526 m  
TypesHolotype: CORBIDI 12931 (Figs. 2–4 in Torres-Carvaljal et al. 2020), adult male, collected on 20 May 2013 by P.J. Venegas, L. Echevarría and M. Gulman.
Paratypes (n = 67): QCAZ, CORBIDI 
DiagnosisDiagnosis: “Macropholidus montanuccii sp. nov. can be distinguished from both M. ataktolepis and M. ruthveni by having a paired series of enlarged middorsal scale rows restricted to the nape (Figs. 2, 3; series continuous to anterior half of body in M. ataktolepis, and continuous to sacral region in M. ruthveni). From M. ataktolepis, M. mon‐ tanuccii sp. nov. further differs in lacking prefrontal scales and having more (33–42, 36.78 ± 1.80) transverse rows of dorsal scales from occipital scale to posterior margin of hind limbs (29–35, 32.8 ± 1.92 in M. ataktolepis). From M. huancabambae, M. montanuccii sp. nov. differs in having shorter and striated dorsal scales (elongate and strongly keeled in M. huancabambae), and more transverse rows of dorsal scales from occipital scale to posterior margin of hind limbs (32–35, 32.2 ± 0.2 in M. huanca‐ bambae). The new species is more similar in morphology to its sister species M. annectens (Fig. 4; character states in parentheses), from which it differs in having fewer transverse rows of dorsal scales from occipital scale to posterior margin of hind limbs (40–48, x = 42.6); fewer—21–28, 23.85 ± 1.77—transverse rows of ventral scales between collar fold and preanals (25–30, x = 27.3); usually a series of black speckles forming a more or less continuous line on middorsum and onto tail (dorsum usually uniform without speckles forming lines; Fig. 4); irregular dark marks on lower lips (lower lips uniform in color); conspicuous ocelli above fore limbs, on neck, and sometimes along flanks extending onto tail (ocelli absent); distinct white dorsolateral stripe from snout to scapular region (white stripe shorter, from snout to nape or to a level above tympanum); and five or more paired, enlarged middorsal scales behind occiput (1–2).” (Torres-Carvajal et al. 2020) 
EtymologyNamed after Richard R. Montanucci, who published a seminal work on the systematics of Pholidobolus lizards in the early 1970’s (Montanucci, 1973) after intensive work along the Andes of Ecuador. Richard Montanucci has dedicated his life to the study of lizards. His work on Pholidobolus lizards is of great importance for anyone interested in gymnophthalmid lizards from the Tropical Andes. 
  • Parra, V., Nunes, P. M. S., & Torres-Carvajal, O. 2020. Systematics of Pholidobolus lizards (Squamata, Gymnophthalmidae) from southern Ecuador, with descriptions of four new species. ZooKeys 954: 109-156
  • Torres-Carvajal O., Venegas P.J., Nunes P.M.S. 2020. Description and phylogeny of a new species of an- dean lizard (Gymnophthalmidae: Cercosaurinae) from the Huancabamba depression. South American Journal of Herpetology 18: 13–23 - get paper here
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